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  • Frequent, repeated defecation by Pteromys volans orii flying squirrels in favored locations results in accumulated droppings. As winter snow melts, embedded droppings dissolve, dripping down to become fertilizer for the relevant tree.
    pteromys-volans-orii-flying-squirrel...tif
  • This Pteromys volans orii flying squirrel is relishing fresh buds of a small branch from a Japanese elm tree (Ulmus davidiana var. japonica).
    pteromys-volans-orii-flying-squirrel...tif
  • This is a marsh tit (Poecile palustris hensoni) hovering beside an icicle formed from the sap of a painted maple tree (Acer pictum). During winter months, small birds like this make use of this calorie-rich food source (essentially frozen maple syrup) to fuel their high metabolisms. The birds fly to an icicle, hover, break off a piece and fly away, all in the blink of eye.
    poecile-palustris-hensoni-marsh-tit-...tif
  • This is a male Pteromys volans orii flying squirrel perched in a safe location high up in the canopy. Behind this knot in the tree was the nest. The squirrel was positioned at the top entrance to the nest, which gave him the ideal vantage point from which to survey the surroundings and keep a lookout for two rival males that were attempting to get the attention of the female inside. Many battles ensued, with much chasing and lots of flying.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel enjoying one of its favorite meals. The animals is eating male catkins of an Alnus japonicus alder tree.
    pteromys-volans-orii-eating-alnus-ja...tif
  • These are male catkins of an East Asian alder tree (Alnus japonica). These catkins are highly prized food among Pteromys volans orii flying squirrels.
    alnus-japonica-alder-male-catkins-ja...tif
  • Pictured here are male catkins of an Alnus japonica elder tree together with female cones. The male catkins comprise many flowers, which produce pollen to pollinate female catkins. Once pollinated, the female catkins develop into woody false cones as seen here. In the autumn the scales of the cones open and disperse their seeds. Male catkins of this species are among the favorite foods of Pteromys volans orii flying squirrels.
    alnus-japonica-alder-catkins-and-con...tif
  • This is a marsh tit (Poecile palustris hensoni) poised to break off a piece of an icicle formed from the sap of a painted maple tree (Acer pictum). During winter months, small birds like this make use of this calorie-rich food source (essentially frozen maple syrup) to fuel their high metabolisms. The birds fly to an icicle, hover, break off a piece and fly away, all in the blink of eye.
    marsh-tit-poecile-palustris-hensoni-...tif
  • This is a long-tailed tit (Aegithalos caudatus) hovering in front of an icicle formed from the sap of a painted maple tree (Acer pictum). During winter months, small birds like this make use of this calorie-rich food source (essentially frozen maple syrup) to fuel their high metabolisms. The birds fly to an icicle, hover, break off a piece and fly away, all in the blink of eye. This is a delicate operation. Sometimes the birds fail to break off a piece, as seen in this series of four photographs. Image 4 in a sequence of 4.
    long-tailed-tit-aegithalos-caudatus-...tif
  • This is a long-tailed tit (Aegithalos caudatus) hovering in front of an icicle formed from the sap of a painted maple tree (Acer pictum). During winter months, small birds like this make use of this calorie-rich food source (essentially frozen maple syrup) to fuel their high metabolisms. The birds fly to an icicle, hover, break off a piece and fly away, all in the blink of eye. This is a delicate operation. Sometimes the birds fail to break off a piece, as seen in this series of four photographs. Image 2 in a sequence of 4.
    long-tailed-tit-aegithalos-caudatus-...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii) peeking out from its nest in the trunk of a tree. Checking the surroundings in this manner prior to emerging at dusk for a night of foraging is common behaviour. Found only in Hokkaido, Japan, these animals prefer small holes to their nests, as this provides a measure of protection from predators like owls and martens. This particular nest entrance is also partially covered by a large white fungus, which obscures it from view. This animal is a sub-species of Siberian flying squirrel and is known locally as ezo-momonga.
    japanese-dwarf-flying-squirrel-at-du...tif
  • A juvenile tufted gray langur sitting in the nook of a tree. This is the Semnopithecus priam thersites subspecies, which is listed as Endangered on the IUCN Red List.
    tufted-gray-langur-juvenile-semnopit...tif
  • This Pteromys volans orii flying squirrel is enjoying a meal of male catkins from an Erman's birch tree (Betula ermanii).
    pteromys-volans-orii-flying-squirrel...tif
  • This is a female whitespotted pygmy filefish (Rudarius ercodes) tending to a clutch of eggs that she has deposited on a cluster of bryozoans that have grown on the branches of a dead tree lodged in the substrate. Development of this species is rapid, with the juveniles hatching in approximately three days.
    rudarius-ercodes-tending-eggs-whites...tif
  • This is a Pteromys volans orii flying squirrel peeking out from its den in a Sakhalin fir tree (Abies sachalinensis) in the soft light of the early winter morning.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a long-tailed tit (Aegithalos caudatus) hovering in front of an icicle formed from the sap of a painted maple tree (Acer pictum). During winter months, small birds like this make use of this calorie-rich food source (essentially frozen maple syrup) to fuel their high metabolisms. The birds fly to an icicle, hover, break off a piece and fly away, all in the blink of eye. Image 3 in a sequence of 3, showing the bird flying away with a piece of the icicle.
    long-tailed-tit-aegithalos-caudatus-...tif
  • This is the Japanese sub-species of Ural owl (Strix uralensis japonica) yawning while sitting in a tree on a snowy day in Hokkaido. These owls are known as Ezo Fukurou in Japanese.
    strix-uralensis-japonica-ural-owl-ya...tif
  • This is a species of flying squirrel called Ezo Momonga in Japanese (Pteromys volans orii). It is a sub-species of Siberian flying squirrels that is found on the island of Hokkaido in Japan. These squirrels are primarily nocturnal and usually shy, though the squirrels can occasionally be seen during the day. They prefer to take up residence in holes made by woodpeckers. Their diet primarily comprises leaves, seeds, cones, buds, sprouts, nuts and berries. Here, the momonga is eating buds of a Japanese elm tree (Ulmus davidiana var. japonica), known as harunire in Japanese. This species does not hibernate.
    ezo-momonga-pteromys-volans-orii-fly...tif
  • Japanese maple trees (Acer sp.), known as momiji in Japanese, produce a riot of colours with the change of seasons. Pictured here are the rich, warm hues of momiji foliage illuminated by the light of morning sun against a clear blue sky.
    japanese-maple-trees-autumn-foliage-...tif
  • Healthy gorgonian sea fans at Deacon's Reef in Milne Bay, Papua New Guinea. Bob Halstead has picked up broken sea fans over many years and planted these sea fans on this wall, producing a beautiful dive site, with overhanging trees visible above.
    red-gorgonian-sea-fan-deacons-reef-m...tif
  • Wall of gorgonian sea fans at Deacon's Reef in Milne Bay, Papua New Guinea. Bob Halstead has picked up broken sea fans over many years and planted these sea fans on this wall, producing a beautiful dive site, with overhanging trees visible above.
    red-gorgonian-sea-fan-deacons-reef-m...tif
  • This is a Pteromys volans orii flying squirrel flying through trees.
    pteromys-volans-orii-flying-squirrel...tif
  • A female Japanese dwarf flying squirrel (Pteromys volans orii) in oestrus often attracts the attention of more than one male. When that happens, males chase one another, leap from and glide among trees, and sometimes engage in physical skirmishes. The male in the background of this image is stalking the male in the foreground in an attempted sneak-attack during one such contest for an eligible female.
    japanese-dwarf-flying-squirrel-males...tif
  • A submerged forest of pneumatophores that supports a vibrant community of mangrove apple (Sonneratia alba) trees in Long Lake, which is a tidal exchange area that connects a saltwater lake inside Ngeruktabel Island with the ocean. The conical pneumatophores pictured here comprise an extended root system that is essential for gas exchange in an anoxic environment. Sonneratia alba communities prefer seaward fringes with high salinity and soft sandy or muddy bottoms.
    Sonneratia-alba-pneumatophore-mangro...tif
  • This is a young Bruguiera gymnorrhiza mangrove plant taking root in the shallow water of Long Lake, part of Ngeruktabel Island in Palau. These mangrove trees create an intricate network of roots, visible behind this young plant, that stablize and lock-in silt and mud, providing a buffer against erosion, as well as serving as shelter for juvenile fish and other marine animals.
    Bruguiera-gymnorrhiza-mangrove-long-...tif
  • This is a Pteromys volans orii flying squirrel en route back to its nest after foraging at the top of 20m+ tall Japanese elm trees (Ulmus davidiana var. japonica) for an extended period of time during snowfall. The squirrel is carrying a branch with fresh elm buds back to its nest—a takeaway meal to relish at home.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel carrying a small branch of Japanese elm (Ulmus davidiana var. japonica) back to its nest. This individual had spent over an hour in the early morning at the top of 20m tall elm trees feasting on fresh buds. This branch is a take-away meal for enjoyment back home.
    pteromys-volans-orii-flying-squirrel...tif
  • Autumn foliage of Momiji Japanese maple trees (Acer sp.) in Kyoto, Japan
    japanese-maple-trees-autumn-foliage-...tif
  • Pictured here are leaves of momiji Japanese maple trees (Acer sp.) that have turned red in autumn. Photographed in Kyoto, Japan.
    japanese-maple-trees-autumn-foliage-...tif
  • This is the moment that one male Japanese dwarf flying squirrel (Pteromys volans orii) ambushed a rival male, sneaking up from behind and leaping on on top of the other male. Male rivalry and competition like this takes place during the reproductive season. The action is rapid, often spanning wide areas as the animals leap and glide among trees.
    japanese-dwarf-flying-squirrel-males...tif
  • This is a male and female pair of Japanese dwarf flying squirrels (Pteromys volans orii), with one inside the nest and the other above. This female (inside the nest) is in oestrus. The male is standing guard. Another male approached repeatedly, resulting in heated confrontation, with the males chasing one another, occasionally leaping and gliding among trees.
    japanese-dwarf-flying-squirrel-matin...tif
  • Colorful momiji Japanese maple trees at Nison-in in Kyoto, Japan.
    momiji-autumn-foliage-nison-in-kyoto...tif
  • Blue Pond in Biei, Hokkaido, Japan in the winter, with snow falling. The pond is manmade, and the trees growing through the water are Japanese larch (Larix kaempferi), deciduous conifers. 北海道の美瑛にある青い池です。
    blue-pond-in-winter-with-snow-hokkai...tif
  • Blue Pond in Biei, Hokkaido, Japan in the winter, with snow falling. The pond is manmade, and the trees growing through the water are Japanese larch (Larix kaempferi), deciduous conifers. 北海道の美瑛にある青い池です。
    blue-pond-in-winter-biei-hokkaido-ja...tif
  • Long Lake, part of Ngeruktabel Island in Palau, is home to a variety of plant life, including a dense forest of mangrove trees. This is a young Bruguiera gymnorrhiza mangrove plant taking root in the shallows. When mature, these mangrove trees create an intricate network of roots that stablize and lock-in silt and mud, providing a buffer against erosion, as well as serving as shelter for juvenile fish and other marine animals.
    bruguiera-gymnorrhiza-young-mangrove...tif
  • This is a Pteromys volans orii flying squirrel taking to the air on a cold, cloudy winter day. Navigating through multiple trees and intersecting branches requires agility and control while airborne.
    pteromys-volans-orii-flying-squirrel...tif
  • Momiji Japanese maple trees displaying an entire rainbow of autumn colors. Photographed at Kita no Tenman-gu in Kyoto, Japan.
    multicolored-momiji-maple-leaves-kit...tif
  • Japanese larch trees (Larix kaempferi) in the blue ice of Blue Pond in Biei, Hokkaido, Japan frozen over during winter.
    japanese-larch-trees-frozen-blue-pon...tif
  • Individual Pteromys volans flying squirrels seem to have favored perches such as this. This individual visited this spot often, sometimes just to sit, sometimes to eat, often as a stopover before gliding to other trees to forage.
    pteromys-volans-orii-flying-squirrel...tif
  • Autumn leaves of Japanese maple trees at Buddhist Tendai sect temple Bishamon-do in Kyoto, Japan
    japanese-maple-trees-autumn-foliage-...tif
  • Chochin paper lantern and Shide (Shinto paper folding) at Kifune Shrine in Kyoto, with brilliant orange-red autumn foliage of momiji Japanese maple trees in the background.
    paper-chochin-lantern-shide-folding-...tif
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Tony Wu Underwater Photography

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