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  • Blue Pond in Biei, Hokkaido, Japan in the winter, with snow falling. The pond is manmade, and the trees growing through the water are Japanese larch (Larix kaempferi), deciduous conifers. 北海道の美瑛にある青い池です。
    blue-pond-in-winter-with-snow-hokkai...tif
  • Blue Pond in Biei, Hokkaido, Japan in the winter, with snow falling. The pond is manmade, and the trees growing through the water are Japanese larch (Larix kaempferi), deciduous conifers. 北海道の美瑛にある青い池です。
    blue-pond-in-winter-biei-hokkaido-ja...tif
  • Japanese larch trees (Larix kaempferi) in the blue ice of Blue Pond in Biei, Hokkaido, Japan frozen over during winter.
    japanese-larch-trees-frozen-blue-pon...tif
  • Waterfall named Fudou no Taki (不動の滝) after the first major snowfall of the winter. Photographed in Biei, Hokkaido, Japan.
    waterfall-in-winter-biei-hokkaido-ja...tif
  • Shirahige no Taki Waterfall (白ひげの滝) in winter. Photographed in Biei, Hokkaido, Japan.
    waterfall-biei-hokkaido-japan-shirah...tif
  • This is a waterfall in Biei, Hokkaido, Japan called Shirahige no Taki (白ひげの滝), photographed in winter after the first snowfall of the season.
    waterfall-biei-hokkaido-shirahige-no...tif
  • This species of crayfish, known as zarigani (Cambaroides japonicus) is endemic to Japan, living in Hokkaido and northern Honshu. They live in areas of low temperature with high water purity. This individual was missing its right front claw.
    zarigani-crayfish-cambaroides-japoni...tif
  • Icicles in the early morning in Hokkaido, Japan
    icicles-pre-dawn-winter-hokkaido-jap...tif
  • Mature male killer whale (Orcinus orca) swimming in the low light of winter in northern Norway
    male-killer-whale-orca-low-light-win...tif
  • This is a pair of Ural owls (Strix uralensis japonica) that have come together at the end of winter in preparation for the spring reproductive season.
    strix-uralensis-japonica-ural-owl-ez...tif
  • This is an Ezo chipmunk (Tamias sibiricus lineatus), recently emerged from winter hibernation.
    tamias-sibiricus-lineatus-ezo-chipmu...tif
  • This is a Pteromys volans orii flying squirrel bathed in the cool light of an early winter morning. These animals are normally nocturnal.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii) flying through the forest on a cold winter day. Usually nocturnal, these animals sometimes become active in the day during reproductive season.
    japanese-flying-squirrel-pteromys-vo...tif
  • Close-up profile of a male Neoclinus bryope chaenopsid blenny during the winter breeding season. This species is known as koke-ginpo in Japanese. Photographed at a magnification of two times life-size.
    neoclinus-bryope-blenny-japan-202101...tif
  • This is a male Neoclinus bryope chaenopsid blenny, active during the winter reproductive season. This species is known as koke-ginpo in Japanese.
    neoclinus-bryope-blenny-male-reprodu...tif
  • This is a long-tailed tit (Aegithalos caudatus) hovering in front of an icicle formed from the sap of a painted maple tree (Acer pictum). During winter months, small birds like this make use of this calorie-rich food source (essentially frozen maple syrup) to fuel their high metabolisms. The birds fly to an icicle, hover, break off a piece and fly away, all in the blink of eye.
    long-tailed-tit-aegithalos-caudatus-...tif
  • This is a long-tailed tit (Aegithalos caudatus) hovering in front of an icicle formed from the sap of a painted maple tree (Acer pictum). During winter months, small birds like this make use of this calorie-rich food source (essentially frozen maple syrup) to fuel their high metabolisms. The birds fly to an icicle, hover, break off a piece and fly away, all in the blink of eye. Image 2 in a sequence of 3, showing the bird breaking a piece off from the tip of the icicle.
    long-tailed-tit-aegithalos-caudatus-...tif
  • This is a male fat greenling (Hexagrammos otakii) protecting eggs. The eggs comprise several clutches from a number of females. During the autumn to winter breeding season, mature males like this one establish territories and adopt a brilliant yellow-orange coloration. The males court passing females, which can choose to spawn with or to ignore a given male.
    hexagrammos-otakii-fat-greenling-mal...tif
  • This is a Japanese red squirrel (Sciurus vulgaris orientis) holding a branch in its mouth. In the final days of winter, this squirrel was building a nest to prepare for the reproductive season.
    sciurus-vulgaris-orientis-ezo-red-sq...tif
  • This is a Pteromys volans orii flying squirrel gliding through the forest early one winter morning.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel gliding through the forest canopy during light snow in winter.
    pteromys-volans-orii-flying-squirrel...tif
  • This is an Ezo chipmunk (Tamias sibiricus lineatus) that has just emerged from hibernation days before this photo was taken. This ground squirrel has lost substantial body mass, as well as the hair on its tail, during the winter.
    tamias-sibiricus-lineatus-ezo-chipmu...tif
  • This is a pair of Pteromys volans orii flying squirrels that have just emerged from their nest early in the morning during the winter reproductive season. The individual on the left is female, right male. After preening together, the pair ascended into the canopy to forage. They mated 10 days after this photo was taken.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a Pteromys volans orii flying squirrel male grooming itself after emerging from his nest on an early winter morning. This male shared the nest with a female. The pair mated 10 days after this photo was taken.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a Pteromys volans orii flying squirrel male grooming itself after emerging from his nest on an early winter morning. This male shared the nest with a female. The pair mated 10 days after this photo was taken.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a long-tailed tit (Aegithalos caudatus) hovering in front of an icicle formed from the sap of a painted maple tree (Acer pictum). During winter months, small birds like this make use of this calorie-rich food source (essentially frozen maple syrup) to fuel their high metabolisms. The birds fly to an icicle, hover, break off a piece and fly away, all in the blink of eye.
    long-tailed-tit-aegithalos-caudatus-...tif
  • This is a male spotty-bellied greenling (Hexagrammos agrammus) protecting several clutches of eggs resulting from spawning with multiple females. During the autumn/ winter breeding season, males keeps watch over developing embryos until they hatch. At the same time, males actively court females that approach, swimming out to greet them and lead them back to carefully prepared spawning areas like the one pictured here. The eyes of developing fish larvae are visible in many of the eggs here, meaning they are close to hatching. Freshly spawned eggs were attached to the opposite side of this cluster. Successful males like this one often attend to a number of egg clusters at different stages of development, each from separate females.
    hexagrammos-agrammus-spotty-bellied-...tif
  • This is a long-tailed tit (Aegithalos caudatus) attempting to take a bite from an icicle formed from the sap of a painted maple tree (Acer pictum). During winter months, small birds like this make use of this calorie-rich food source (essentially frozen maple syrup) to fuel their high metabolisms. The birds often need to hover in front of icicles, but in this instance, there was a convenient perch. In the end, the icicle proved too thick for the bird to be able break off a piece with its small mouth.
    long-tailed-tit-aegithalos-caudatus-...tif
  • This is a north Atlantic humpback whale (Megaptera novaeangliae novaeangliae) pursuing a large school of herring in the cold, dark water of Arctic winter. The humpback whale swallowed several mouthfuls of fish during this foraging period.
    humpback-whale-pursuing-herring-norw...tif
  • This is a long-tailed tit (Aegithalos caudatus) hovering in front of an icicle formed from the sap of a painted maple tree (Acer pictum). During winter months, small birds like this make use of this calorie-rich food source (essentially frozen maple syrup) to fuel their high metabolisms. The birds fly to an icicle, hover, break off a piece and fly away, all in the blink of eye. A piece of the icicle is visible in the bird’s mouth.
    long-tailed-tit-aegithalos-caudatus-...tif
  • This is a 40cm mature male smooth lumpsucker (Aptocyclus ventricosus) watching over a brood of eggs. This species resides in deep water, down to about 1700m. During the winter to spring breeding season, mature fish come up to shallow water. After fertilizing a brood of eggs laid by a female fish, the male watches over and protects the developing young. Adult fish die during the breeding season, the female after laying eggs, the male once the juveniles have hatched. Following extended observation, I realized that the eggs in this photo were not viable. They had not been fertilized and were therefore never going to produce juveniles. The male, unaware of this fact, was destined continue to protect the infertile eggs, weakening day by day, eventually succumbing to death.
    aptocyclus-ventricosus-smooth-lumpsu...tif
  • Two adult southern hemisphere humpback whales (Megaptera novaeangliae australis) tail slapping together. This pair was traveling north together in the winter. The one on the left is female; the one on the right male.
    southern-humpback-whale-megaptera-no...tif
  • Red fox, Sakhalin subspecies (Vulpes vulpes schrencki) foraging for food in the winter snow of Hokkaido. Photographed in Shiretoko National Park.
    red-fox-sakhalin-subspecies-vulpes-v...tif
  • Female Japanese sika deer (Cervus nippon yesoensis) with young fawn. The deer were foraging for food during the winter in Utoro, Hokkaido, Japan.
    japanese-sika-deer-Cervus-nippon-yes...tif
  • Feces from an injured humpback whale (Megaptera novaeangliae) encountered during the winter season in the waters around Vava'u, Kingdom of Tonga. The humpback whale was entangled in a series of longlines and hooks and most likely died. Bycatch is a serious problem associated with longline fishing. Even whales occasionally fall victim to this indiscriminate fishing practice.
    humpback-whale-feces-defecation-tong...tif
  • This is a Japanese red squirrel (Sciurus vulgaris orientis) leaping among the branches of a fir tree while collecting material for the construction of a nest in the final days of winter. The squirrel was preparing for the upcoming reproductive season.
    sciurus-vulgaris-orientis-ezo-red-sq...tif
  • This is a Pteromys volans orii flying squirrel sitting in a favorite spot during an overcast winter morning.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel pictured in the split-second before landing on a tree. The Sakhalin fir trees in the background are nearly completely covered in snow from a large winter storm.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a marsh tit (Poecile palustris hensoni) hovering beside an icicle formed from the sap of a painted maple tree (Acer pictum). During winter months, small birds like this make use of this calorie-rich food source (essentially frozen maple syrup) to fuel their high metabolisms. The birds fly to an icicle, hover, break off a piece and fly away, all in the blink of eye.
    poecile-palustris-hensoni-marsh-tit-...tif
  • This black-headed gull (Larus ridibundus) has just dived down to steal a fish (possibly juvenile Stichaeus grigorjewi) from an unfortunate common merganser (Mergus merganser). The gull is in winter plumage.
    larus-ridibundus-gull-stealing-fish-...tif
  • This is a Pteromys volans orii flying squirrel eating one of its favorite foods—catkins from a Japanese alder tree (Alnus japonica). This individual had descended to ground level during a winter evening to forage for fallen catkins. Descending in this manner is unusual behavior.
    pteromys-volans-orii-flying-squirrel...tif
  • This is an Ezo chipmunk (Tamias sibiricus lineatus) that has just emerged from hibernation days before this photo was taken. This ground squirrel has lost substantial body mass, as well as the hair on its tail, during the winter.
    tamias-sibiricus-lineatus-ezo-chipmu...tif
  • This is a Pteromys volans orii flying squirrel male pausing after grooming himself during an early winter morning. This male shared the nest with a female, which joined him shortly after this photo was taken. The pair mated 10 days after this photo was taken.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a pair of Pteromys volans orii flying squirrels that have just emerged from their nest early in the morning during the winter reproductive season. The individual on the left is female, right male. After preening together, the pair ascended into the canopy to forage. They mated 10 days after this photo was taken.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a Pteromys volans orii flying squirrel male grooming itself after emerging from his nest on an early winter morning. This male shared the nest with a female. The pair mated 10 days after this photo was taken.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a pair of Japanese dwarf flying squirrels (Pteromys volans orii). The one in front is male. The one peeking out from behind is female. They are situated at the top entrance to their nest, caressed by warm light from the setting winter sun. This pair mated several times throughout the day. There was another male in the area, a challenger. The female mated with the other male as well, when this one was not looking.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a female East Asian common octopus (Octopus sinensis) protecting a clutch of eggs during the winter, which is not the usual time of year for this species to brood eggs. Reproductive activity is most common during the summer months. This female was eventually successful. Some eggs started to hatch after 62 days, running until 76 days. Hatching during the normal season requires only a few days.
    octopus-sinensis-east-asian-common-o...tif
  • This is a well-camouflaged male Neoclinus bryope chaenopsid blenny peeking out from the safety of his hole in the reef, on the lookout for eligible females during the winter reproductive season. This species is known as koke-ginpo in Japanese.
    neoclinus-bryope-blenny-male-reprodu...tif
  • This is a male Neoclinus bryope chaenopsid blenny active during the winter reproductive season. This species is known as koke-ginpo in Japanese.
    neoclinus-bryope-blenny-male-reprodu...tif
  • This is a long-tailed tit (Aegithalos caudatus) with winter plumage. Photographed in Hokkaido, Japan.
    long-tailed-tit-aegithalos-caudatus-...tif
  • This is a long-tailed tit (Aegithalos caudatus) with winter plumage. Photographed in Hokkaido, Japan.
    long-tailed-tit-aegithalos-caudatus-...tif
  • Juvenile 10cm Chinese pond turtle (Mauremys reevesii) sitting comfortably in a plumbing joint in front of a space heater during winter. This species is semiaquatic in the wild, found in marshes, ponds, streams and similar bodies of shallow water. It is listed as Endangered on the IUCN Red List, threatened by several causes, including competition from introduced species, loss of habitat, and use in Chinese medicine. This species is also popular in the global pet trade. This individual was found on a road in Japan, far from water, when it was only 2.8cm, perhaps picked up and transported by a crow shortly after birth. Though the species had earlier been considered native to Japan, genetic testing in recent years suggests multiple introductions from outside Japan.
    mauremys-reevesii-chinese-pond-turtl...tif
  • This is a marsh tit (Poecile palustris hensoni) poised to break off a piece of an icicle formed from the sap of a painted maple tree (Acer pictum). During winter months, small birds like this make use of this calorie-rich food source (essentially frozen maple syrup) to fuel their high metabolisms. The birds fly to an icicle, hover, break off a piece and fly away, all in the blink of eye.
    marsh-tit-poecile-palustris-hensoni-...tif
  • This is a long-tailed tit (Aegithalos caudatus) hovering in front of an icicle formed from the sap of a painted maple tree (Acer pictum). During winter months, small birds like this make use of this calorie-rich food source (essentially frozen maple syrup) to fuel their high metabolisms. The birds fly to an icicle, hover, break off a piece and fly away, all in the blink of eye. This is a delicate operation. Sometimes the birds fail to break off a piece, as seen in this series of four photographs. Image 3 in a sequence of 4.
    long-tailed-tit-aegithalos-caudatus-...tif
  • This is a long-tailed tit (Aegithalos caudatus) hovering in front of an icicle formed from the sap of a painted maple tree (Acer pictum). During winter months, small birds like this make use of this calorie-rich food source (essentially frozen maple syrup) to fuel their high metabolisms. The birds fly to an icicle, hover, break off a piece and fly away, all in the blink of eye. This is a delicate operation. Sometimes the birds fail to break off a piece, as seen in this series of four photographs. Image 1 in a sequence of 4.
    long-tailed-tit-aegithalos-caudatus-...tif
  • This is a long-tailed tit (Aegithalos caudatus) hovering in front of an icicle formed from the sap of a painted maple tree (Acer pictum). During winter months, small birds like this make use of this calorie-rich food source (essentially frozen maple syrup) to fuel their high metabolisms. The birds fly to an icicle, hover, break off a piece and fly away, all in the blink of eye. Image 1 in a sequence of 3, showing the bird grab the icicle.
    long-tailed-tit-aegithalos-caudatus-...tif
  • This is a long-tailed tit (Aegithalos caudatus) hovering under an icicle formed from the sap of a painted maple tree (Acer pictum). During winter months, small birds like this make use of this calorie-rich food source (essentially frozen maple syrup) to fuel their high metabolisms. The birds fly to an icicle, hover, break off a piece and fly away, all in the blink of eye.
    long-tailed-tit-aegithalos-caudatus-...tif
  • This is a male fat greenling (Hexagrammos otakii) tending to eggs. The eggs comprise several clutches from a number of females. During the autumn to winter breeding season, mature males like this one establish territories and adopt a brilliant yellow-orange coloration. The males court passing females, which can choose to spawn with or to ignore a given male.
    hexagrammos-otakii-fat-greenling-mal...tif
  • This is a species of flying squirrel called Ezo Momonga in Japanese (Pteromys volans orii). It is a sub-species of Siberian flying squirrels that is found on the island of Hokkaido in Japan. These squirrels are primarily nocturnal and usually shy, though the squirrels can occasionally be seen during the day. They prefer to take up residence in holes made by woodpeckers. Their diet primarily comprises leaves, seeds, cones, buds, sprouts, nuts and berries. This individual paused in this position for a few moments after emerging one winter evening from its hole in a Sakhalin fir tree (Abies sachalinensis), known as todomatsu in Japanese. This species does not hibernate.
    ezo-momonga-pteromys-volans-orii-fly...tif
  • Atlantic puffin (Fratercula arctica) basking in the warm light of winter sunrise, Iceland
    atlantic-puffin-fratercula-arctica-i...tif
  • Red fox, Sakhalin subspecies (Vulpes vulpes schrencki) foraging for food in the winter snow of Hokkaido. Photographed in Shiretoko National Park.
    red-fox-sakhalin-subspecies-vulpes-v...tif
  • Japanese spotted deer (Cervus nippon yesoensis) with young fawn and two more deer in the background. The deer were foraging for food during winter. Photographed in Shiretoko National Park, Utoro, Hokkaido, Japan.
    japanese-spotted-deer-cervus-nippon-...tif
  • This is a Pteromys volans orii flying squirrel enjoying a good scratch early one winter morning.
    pteromys-volans-orii-flying-squirrel...tif
  • Frequent, repeated defecation by Pteromys volans orii flying squirrels in favored locations results in accumulated droppings. As winter snow melts, embedded droppings dissolve, dripping down to become fertilizer for the relevant tree.
    pteromys-volans-orii-flying-squirrel...tif
  • Pictured here is a pair of Ural owls (Strix uralensis japonica) in their nest during late afternoon on a winter day. The smaller individual on the lower left is the male. This pair was preparing to have chicks.
    strix-uralensis-japonica-ural-owl-ez...tif
  • This is a pair of Ural owls (Strix uralensis japonica) that have come together at the end of winter in preparation for the spring reproductive season.
    strix-uralensis-japonica-ural-owl-ez...tif
  • This is a Pteromys volans orii flying squirrel taking to the air on a cold, cloudy winter day. Navigating through multiple trees and intersecting branches requires agility and control while airborne.
    pteromys-volans-orii-flying-squirrel...tif
  • This is an Ezo chipmunk (Tamias sibiricus lineatus) that has just emerged from hibernation days before this photo was taken. During the winter, the chipmunk lost substantial body mass, as well as the hair on its tail.
    tamias-sibiricus-lineatus-ezo-chipmu...tif
  • This is an Ezo chipmunk (Tamias sibiricus lineatus) foraging for food. It had just emerged from hibernation days before this photo was taken. This ground squirrel has lost substantial body mass, as well as the hair on its tail, during the winter.
    tamias-sibiricus-lineatus-ezo-chipmu...tif
  • This is a pair of mating Pteromys volans orii flying squirrels engaged in copulation on a cold winter day.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a pair of Pteromys volans orii flying squirrels that have just emerged from their nest early in the morning during the winter reproductive season. The individual facing the camera is male. After preening together, the pair ascended into the canopy to forage. They mated 10 days after this photo was taken.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a pair of Pteromys volans orii flying squirrels that have just emerged from their nest early in the morning during the winter reproductive season. The individual on the left is female, right male. After preening together, the pair ascended into the canopy to forage. They mated 10 days after this photo was taken.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a Pteromys volans orii flying squirrel male grooming itself after emerging from his nest on an early winter morning. This male shared the nest with a female. The pair mated 10 days after this photo was taken.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a portrait of a Pteromys volans orii flying squirrel male illuminated by the light of an early winter morning. He had just emerged from his nest, followed shortly thereafter by a female. The pair mated 10 days after this photo was taken.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a Pteromys volans orii flying squirrel peeking out from its den in a Sakhalin fir tree (Abies sachalinensis) in the soft light of the early winter morning.
    japanese-flying-squirrel-pteromys-vo...tif
  • Known as koke-ginpo in Japanese, this chaenopsid blenny is a male Neoclinus bryope. This dark coloration with the bright blue dorsal fin spot is indicative of the winter breeding season in this area, where he and other males shake, shimmy and perform to attract the attention of females. Photographed at a magnification of two times life-size.
    neoclinus-bryope-blenny-male-reprodu...tif
  • This is a long-tailed tit (Aegithalos caudatus) with winter plumage. Photographed in Hokkaido, Japan.
    long-tailed-tit-aegithalos-caudatus-...tif
  • This is a long-tailed tit (Aegithalos caudatus) hovering in front of an icicle formed from the sap of a painted maple tree (Acer pictum). During winter months, small birds like this make use of this calorie-rich food source (essentially frozen maple syrup) to fuel their high metabolisms. The birds fly to an icicle, hover, break off a piece and fly away, all in the blink of eye. This is a delicate operation. Sometimes the birds fail to break off a piece, as seen in this series of four photographs. Image 4 in a sequence of 4.
    long-tailed-tit-aegithalos-caudatus-...tif
  • This is a long-tailed tit (Aegithalos caudatus) hovering in front of an icicle formed from the sap of a painted maple tree (Acer pictum). During winter months, small birds like this make use of this calorie-rich food source (essentially frozen maple syrup) to fuel their high metabolisms. The birds fly to an icicle, hover, break off a piece and fly away, all in the blink of eye. This is a delicate operation. Sometimes the birds fail to break off a piece, as seen in this series of four photographs. Image 2 in a sequence of 4.
    long-tailed-tit-aegithalos-caudatus-...tif
  • This is a long-tailed tit (Aegithalos caudatus) hovering in front of an icicle formed from the sap of a painted maple tree (Acer pictum). During winter months, small birds like this make use of this calorie-rich food source (essentially frozen maple syrup) to fuel their high metabolisms. The birds fly to an icicle, hover, break off a piece and fly away, all in the blink of eye. Image 3 in a sequence of 3, showing the bird flying away with a piece of the icicle.
    long-tailed-tit-aegithalos-caudatus-...tif
  • This is a long-tailed tit (Aegithalos caudatus) hovering in front of an icicle formed from the sap of a painted maple tree (Acer pictum). During winter months, small birds like this make use of this calorie-rich food source (essentially frozen maple syrup) to fuel their high metabolisms. The birds fly to an icicle, hover, break off a piece and fly away, all in the blink of eye.
    long-tailed-tit-aegithalos-caudatus-...tif
  • This is a male fat greenling (Hexagrammos otakii) protecting eggs. The eggs comprise several clutches from a number of females. During the autumn to winter breeding season, mature males like this one establish territories and adopt a brilliant yellow-orange coloration. The males court passing females, which can choose to spawn with or to ignore a given male.
    hexagrammos-otakii-fat-greenling-mal...tif
  • This is a male fat greenling (Hexagrammos otakii) tending to eggs. The eggs comprise several clutches from a number of females.  During the autumn to winter breeding season, mature males like this one establish territories and adopt a brilliant yellow-orange coloration. The males court passing females, which can choose to spawn with or to ignore a given male.
    hexagrammos-otakii-fat-greenling-mal...tif
  • Mature bull killer whale (Orcinus orca) in the cold, dark winter waters of northern Norway. Note the rake marks on the orca’s body, perhaps from encounters with other male orcas.
    mature-male-killer-whale-orcinus-orc...tif
  • Japanese spotted deer (Cervus nippon yesoensis) foraging for food during winter, along the edge of a slope overlooking the Okhotsk Sea. Photographed in Shiretoko National Park, Utoro, Hokkaido, Japan.
    japanese-spotted-deer-portrait-cervu...tif
  • Juvenile Sika Deer (Cervus nippon yesoensis) foraging for food in the winter. Here the deer is nibbling on Japanese rose (Rosa rugosa). This is the Hokkaido subspecies of Cervus nippon. Photographed in Biei, Hokkaido, Japan. エゾシカ, 美瑛、北海道
    japanese-sika-deer-hokkaido-subspeci...tif
  • Male humpback whale (Megaptera novaeangliae) defecating while socializing with two other male humpback whales. Photographed during the winter breeding season in the Kingdom of Tonga.
    humpback-whale-defecation-tonga-2010...tif
  • Feces from an injured humpback whale (Megaptera novaeangliae) encountered during the winter season in the waters around Vava'u, Kingdom of Tonga. The humpback whale was entangled in a series of longlines and hooks and most likely died. Bycatch is a serious problem associated with longline fishing. Even whales occasionally fall victim to this indiscriminate fishing practice.
    humpback-whale-feces-defecation-tong...tif
  • Whooper swan (Cygnus cygnus) visiting wintering grounds in Hokkaido, Japan
    cygnus-cygnus-whooper-swan-japan-202...tif
  • Whooper swans (Cygnus cygnus) visiting wintering grounds in Hokkaido, Japan
    cygnus-cygnus-whooper-swans-japan-20...tif
  • Whooper swan (Cygnus cygnus) visiting wintering grounds in Hokkaido, Japan. The swan is turned tail-end up while foraging for food.
    cygnus-cygnus-whooper-swan-foraging-...tif
  • Whooper swan (Cygnus cygnus) visiting wintering grounds in Hokkaido, Japan
    cygnus-cygnus-whooper-swan-japan-202...tif
  • This is a whooper swan (Cygnus cygnus) in flight. Many of these birds spend their winters in Hokkaido, Japan, then fly north again with the arrival of warmer weather in spring.
    cygnus-cygnus-whooper-swan-japan-202...tif
  • Whooper swan (Cygnus cygnus) in Hokkaido wintering grounds eating seagrass that it grabbed while wading in a shallow bay
    cygnus-cygnus-whooper-swan-japan-202...tif
  • A pair of whooper swans (Cygnus cygnus) visiting wintering grounds in Hokkaido, Japan
    cygnus-cygnus-whooper-swans-japan-20...tif
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Tony Wu Underwater Photography

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