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  • Pictured here is a pair of a Japanese dwarf flying squirrel (Pteromys volans orii), one peeking out from the nest, and another perched on a branch above. These squirrels are preparing for a night of foraging, having emerged shortly after sunset. Known locally as ezo-momonga, this sub-species is a  sub-species of Siberian flying squirrel and is found only in Hokkaido, Japan. It is primarily nocturnal. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range.
    japanese-dwarf-flying-squirrels-at-d...tif
  • This is a pair of Japanese dwarf flying squirrels (Pteromys volans orii) engaged in copulation. This is an extended process spanning many hours, during which the male often needs to fend off rivals while engaging in repeated bouts of copulation.
    japanese-dwarf-flying-squirrels-mati...tif
  • This is a pair of Japanese dwarf flying squirrels (Pteromys volans orii) engaged in copulation. This is an extended process spanning many hours, during which the male often needs to fend off rivals while engaging in repeated bouts of copulation.
    japanese-dwarf-flying-squirrels-mati...tif
  • This series of six images shows a pair of Japanese dwarf flying squirrels (Pteromys volans orii) that have just emerged from a nest in the tree. The male pursues the female; the pair copulate; and the female returns to the nest. Over the course of a day or so, the male copulates as many times as possible. Rival males sometimes approach, which can result in a heated battle between males.
    japanese-dwarf-flying-squirrels-copu...tif
  • This is a pair of Japanese dwarf flying squirrels (Pteromys volans orii) engaged in copulation. This is an extended process spanning many hours, during which the male often needs to fend off rivals while engaging in repeated bouts of copulation.
    japanese-dwarf-flying-squirrels-mati...tif
  • This is a pair of Japanese dwarf flying squirrels (Pteromys volans orii) engaged in copulation. This is an extended process spanning many hours, during which the male often needs to fend off rivals while engaging in repeated bouts of copulation.
    japanese-dwarf-flying-squirrels-mati...tif
  • This is a pair of Japanese dwarf flying squirrels (Pteromys volans orii) engaged in copulation. This is an extended process spanning many hours, during which the male often needs to fend off rivals while engaging in repeated bouts of copulation. As shown here, the female can, and often does, put up some measure of resistance, occasionally dislodging the male completely. The male’s penis is visible.
    japanese-dwarf-flying-squirrels-mati...tif
  • This is a pair of Japanese dwarf flying squirrels (Pteromys volans orii) engaged in copulation. This is an extended process spanning many hours, during which the male often needs to fend off rivals while engaging in repeated bouts of copulation.
    japanese-dwarf-flying-squirrels-mati...tif
  • This is a pair of Japanese dwarf flying squirrels (Pteromys volans orii) engaged in copulation. This is an extended process spanning many hours, during which the male often needs to fend off rivals while engaging in repeated bouts of copulation. As shown here, the female can, and often does, put up some measure of resistance, occasionally dislodging the male completely. The male’s penis is visible.
    japanese-dwarf-flying-squirrels-mati...tif
  • This is a pair of Japanese dwarf flying squirrels (Pteromys volans orii), photographed during breeding season. The individual higher up on the tree is a female in oestrus. In the first two images in this sequence of three, the male approaches the female from behind to attempt copulation. In the third image, the female spins around to indicate in no uncertain terms that she is not ready. The pair’s nest is in the crack in the tree.
    japanese-dwarf-flying-squirrels-copu...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii), a sub-species of Siberian flying squirrel. Known locally as ezo-momonga, this sub-species is found only in Hokkaido, Japan. It is primarily nocturnal. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range. This individual has just emerged shortly after sunset from its nest, a cavity in the tree pictured. After preening for a bit, the flying squirrel ascended into the canopy to forage for the night. While these squirrels are known to nest in a number of tree species, they appear to prefer Sakhalin fir trees (Abies sachalinensis).
    japanese-dwarf-flying-squirrel-ptero...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii), a sub-species of Siberian flying squirrel. Known locally as ezo-momonga, this sub-species is found only in Hokkaido, Japan. It is primarily nocturnal. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range. This individual has just emerged shortly after sunset from its nest, a cavity in the tree pictured. After preening for a bit, the flying squirrel ascended into the canopy to forage for the night. While these squirrels are known to nest in a number of tree species, they appear to prefer Sakhalin fir trees (Abies sachalinensis).
    japanese-dwarf-flying-squirrel-at-du...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-20200...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous. The squirrel’s paddle-like tail is visible from this angle, flopped over its back. This animal uses its tail somewhat like a rudder during flight, helping to adjust direction.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous. The squirrel paused for a moment to groom himself.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous. The squirrel took a moment here to scratch his back.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii), a sub-species of Siberian flying squirrel. Known locally as ezo-momonga, this sub-species is found only in Hokkaido, Japan. It is primarily nocturnal. This indiividual has just emerged from its nest at dusk, and is taking a moment before heading out for a night of foraging. From this perspective, it is possible to see how the squirrel grasps and maintains a grip on delicate branches. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range.
    japanese-dwarf-flying-squirrel-at-du...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii), a sub-species of Siberian flying squirrel. getting ready to leave its nest to set out for a night of foraging. Known locally as ezo-momonga, this sub-species is found only in Hokkaido, Japan. It is primarily nocturnal. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range.
    japanese-dwarf-flying-squirrel-at-du...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii) flying through the forest on a cold winter day. Usually nocturnal, these animals sometimes become active in the day during reproductive season.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This male was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This male was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This male was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-20200...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii) sitting on a branch. This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This male was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This male was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii), a sub-species of Siberian flying squirrel, emerging from its nest at dusk to set out for a night of foraging. Known locally as ezo-momonga, this sub-species is found only in Hokkaido, Japan. It is primarily nocturnal. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range.
    japanese-dwarf-flying-squirrel-at-du...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii), a sub-species of Siberian flying squirrel. that has just emerged from its nest before setting out for a night of foraging. Known locally as ezo-momonga, this sub-species is found only in Hokkaido, Japan. It is primarily nocturnal. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range.
    japanese-dwarf-flying-squirrel-at-du...tif
  • This Japanese dwarf flying squirrel (Pteromys volans orii) is just emerging from its nest in the trunk of a tree at dusk to prepare for a night of foraging. Found only in Hokkaido, Japan, these animals prefer small holes to their nests, as this provides a measure of protection from predators like owls and martens. This particular nest entrance is also partially covered by a large white fungus, which obscures it from view. This animal is a sub-species of Siberian flying squirrel and is known locally as ezo-momonga.
    japanese-dwarf-flying-squirrel-emerg...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii), gliding from the top of one tree to another. Known locally as ezo-momonga, this sub-species of Siberian flying squirrel is found only in Hokkaido, Japan. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range. Although these animals are primarily nocturnal, they sometimes emerge during the day. Usually this is for brief durations to urinate or defecate. In this instance, this particular individual was one of two males competing for the right to mate with a female. Such reproduction-related competition sometimes takes place during daylight hours.
    japanese-dwarf-flying-squirrel-airbo...tif
  • This is a Pteromys volans orii flying squirrel flying through trees.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a male Pteromys volans orii flying squirrel perched in a safe location high up in the canopy. Behind this knot in the tree was the nest. The squirrel was positioned at the top entrance to the nest, which gave him the ideal vantage point from which to survey the surroundings and keep a lookout for two rival males that were attempting to get the attention of the female inside. Many battles ensued, with much chasing and lots of flying.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This male was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii) peeking out from its nest in the trunk of a tree. Checking the surroundings in this manner prior to emerging at dusk for a night of foraging is common behaviour. Found only in Hokkaido, Japan, these animals prefer small holes to their nests, as this provides a measure of protection from predators like owls and martens. This particular nest entrance is also partially covered by a large white fungus, which obscures it from view. This animal is a sub-species of Siberian flying squirrel and is known locally as ezo-momonga.
    japanese-dwarf-flying-squirrel-at-du...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii), looking out from his nest. This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This male was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii), a sub-species of Siberian flying squirrel, peeking out from its nest before emerging for a night of foraging. Known locally as ezo-momonga, this sub-species is found only in Hokkaido, Japan. It is primarily nocturnal. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range.
    japanese-dwarf-flying-squirrel-at-du...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii) taking a moment to gather itself after emerging from its nest to head out for a night of foraging. Known locally as ezo-momonga, this sub-species of Siberian flying squirrel is found only in Hokkaido, Japan. It is primarily nocturnal. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range.
    japanese-dwarf-flying-squirrel-at-du...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii) that has just come out from its nest., entrance visible to left and above the animal’s head. These flying squirrels are nocturnal, but occasionally emerge during daylight hours.
    japanese-dwarf-flying-squirrel-20200...tif
  • When a Japanese dwarf flying squirrel (Pteromys volans orii) emerges from its nest at dusk, one of the first orders of business is often to relieve itself. The individual pictured here is defecating (pellets visible in the appropriate location) while grasping a small twig for stability. Flying squirrel droppings are used in traditional Chinese herbal medicine for ailments related to blood circulation and abdominal pain. The droppings are known as Wu Ling Zhi.
    japanese-dwarf-flying-squirrel-defec...tif
  • This is a Pteromys volans orii flying squirrel flying directly overhead.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii Japanese dwarf flying squirrel flying through the forest during a flurry of activity involving five individuals, two females and three males. The two females were in estrus.
    japanese-flying-squirrel-pteromys-vo...tif
  • Shown here is the split-second when a Pteromys volans orii flying squirrel makes contact with a tree upon landing. The squirrel first uses its hind limbs to gain a foothold, then secures its front limbs. The squirrel had been foraging during light snowfall.
    pteromys-volans-orii-flying-squirrel...tif
  • This is an unusual behavior. This Pteromys volans orii flying squirrel had stashed food in an unused bird's nest at some point. On this overcast day, the squirrel plucked out and consumed male catkins of an Erman's birch (Betula ermanii), one after another, until the food stash was exhausted.
    pteromys-volans-orii-flying-squirrel...tif
  • The frustrating moment when a Pteromys orii volans flying squirrel passed directly over my head from behind. I spun around and pressed the shutter on instinct from waist height as the squirrel flew past mere centimeters above my head.
    pteromys-volans-orii-flying-squirrel...tif
  • The Pteromys volans orii flying squirrel male seen fleeing up the tree in this photograph had just been chased off by the male poking his head out of the nest. The reason for this conflict was a female inside the nest. She was approaching estrus.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel gliding through the forest in the early morning.
    pteromys-volans-orii-flying-squirrel...tif
  • This Pteromys volans orii flying squirrel had just pulled its face out of the snow. It had been digging in search of food that it had previously gathered and buried for future consumption.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel consuming male catkins from an Erman's birch tree (Betula ermanii).
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel gliding through the forest.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel en route back to its nest after foraging at the top of 20m+ tall Japanese elm trees (Ulmus davidiana var. japonica) for an extended period of time during snowfall. The squirrel is carrying a branch with fresh elm buds back to its nest—a takeaway meal to relish at home.
    pteromys-volans-orii-flying-squirrel...tif
  • This Pteromys volans orii flying squirrel is eating fresh needles of a Sakhalin fir tree (Abies sachalinensis) during a light snowfall. One of the squirrel's teeth is visible from this angle.
    pteromys-volans-orii-flying-squirrel...tif
  • Depicted here is the split-second prior to this Pteromys volans orii flying squirrel landing on a tree. The squirrel's limbs are stretched forward to brace for impact and to ensure that the animal can move immediately when it makes contact.
    pteromys-volans-orii-flying-squirrel...tif
  • This Japanese flying squirrel (Pteromys volans orii) has just emerged from its nest at dusk, before setting out for a night of a foraging in the tree tops.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel bathed in the cool light of an early winter morning. These animals are normally nocturnal.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel defecating, having assumed a comfortable seated position at the entrance of its nest for the purpose. It is clear from the dark trail below the droppings that the residents of this nest also urinate here. This is unusual. In most cases, these squirrels relieve themselves outside, some distance from their nests.
    pteromys-volans-orii-flying-squirrel...tif
  • Pteromys volans orii flying squirrels often have preferred places for defecation. Finding droppings like this is a sign that there are flying squirrels nearby. Dropping piles this large are unusual.
    pteromys-volans-orii-defecation-pano...tif
  • This is a Pteromys volans orii Japanese dwarf flying squirrel gliding across the top of the tree canopy after sunset. The moon is visible, six days prior to being full. This individual was part of a ruckus of reproductive season activity stretching over many hours that involved two females and three males.
    japanese-flying-squirrel-pteromys-vo...tif
  • Japanese dwarf flying squirrel (Pteromys volans orii) perched at the juncture of a small branch and tree trunk. This species is normally nocturnal. This one appeared during briefly during the day, but soon returned to its nest.
    japanese-dwarf-flying-squirrel-20200...tif
  • This is a Pteromys volans orii flying squirrel gliding through the forest early one winter morning.
    pteromys-volans-orii-flying-squirrel...tif
  • This Pteromys volans orii flying squirrel has carried a few sticks of Japanese elm (Ulmus davidiana var. japonica) back to its nest at the end of foraging to enjoy a takeaway meal before going inside.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel peeking out from the entrance to its nest just before a major snowstorm hit. The northwest side of the tree was already coated with snow, and the winds had picked up considerably. The squirrel decided not to emerge at the time.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a female Pteromys volans orii flying squirrel feasting on a branch that she cut from a nearby Sakhalin fir tree (Abies sachalinensis).
    pteromys-volans-orii-flying-squirrel...tif
  • This Pteromys volans orii flying squirrel has just started to consume male catkins from an Erman's birch tree (Betula ermanii).
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel enjoying a meal of Japanese elm buds (Ulmus davidiana var. japonica) while perched more than 20m above the ground. Note the squirrel's perfect balance while seated at the end of tiny branches.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel enjoying an early morning scratch.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel making its final approach back to the nest after foraging. The animal blends in perfectly with its environment, a snow-covered forest. The entrance to the nest is a narrow crack in the Erman's birch (Betula ermanii) tree.
    pteromys-volans-orii-flying-squirrel...tif
  • This composite image shows the landing sequence of a Pteromys volans orii flying squirrel, covering a distance of about 4m in less than half a second. This implies an average flight speed in the range of 30-35kph. The squirrel launched from a higher point on a nearby tree, gliding downward to cover distance, then pulling up as shown here to land on the targeted tree. Note that the distance traveled between frames decreases as the squirrel approaches the tree, indicating deceleration as the animal shifts from horizontal to vertical attitude.
    pteromys-volans-orii-flying-squirrel...tif
  • This is an unusual behavior. This Pteromys volans orii flying squirrel had stashed food in an unused bird's nest at some point. On this overcast day, the squirrel plucked out and consumed male catkins of an Erman's birch (Betula ermanii), one after another, until the food stash was exhausted.
    pteromys-volans-orii-flying-squirrel...tif
  • This is an unusual behavior. This Pteromys volans orii flying squirrel had stashed food in an unused bird's nest at some point. On this overcast day, the squirrel plucked out and consumed male catkins of an Erman's birch (Betula ermanii), one after another, until the food stash was exhausted.
    pteromys-volans-orii-flying-squirrel...tif
  • Depicted here is the split-second prior to this Pteromys volans orii flying squirrel landing on a tree. The squirrel has adjusted its attitude in mid-flight toward a relatively vertical position to reduce speed. It has also folded its wings inward and stretched its limbs forward in preparation for grasping the tree ahead.
    pteromys-volans-orii-flying-squirrel...tif
  • This Japanese flying squirrel (Pteromys volans orii) is perched on a comfortable, safe location high above the ground, where it is eating one of its favorite foods, the buds of a Japanese elm tree (Ulmus davidiana var. japonica). Normally nocturnal, these squirrels are sometimes active in the early hours of daylight.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel emerging from a hideaway at the base of a tall tree. This individual was just digging its way out after snowfall had accumulated and blocked the entrance.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel defecating, having assumed a comfortable seated position at the entrance of its nest for the purpose. It is clear from the dark trail below the droppings that the residents of this nest also urinate here. This is unusual. In most cases, these squirrels relieve themselves outside, some distance from their nests.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel male grooming itself after emerging from his nest on an early winter morning. This male shared the nest with a female. The pair mated 10 days after this photo was taken.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a Pteromys volans orii flying squirrel male grooming itself after emerging from his nest on an early winter morning. This male shared the nest with a female. The pair mated 10 days after this photo was taken.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii) that has found a comfortable place to perch on an overcast day after emerging from its nest. These animals line the inside of their nests with soft materials for comfort and warmth. Some of this nest lining is visible on this squirrel’s head.
    japanese-dwarf-flying-squirrel-20200...tif
  • In the midst of competition for the right to mate with a female Japanese dwarf flying squirrel (Pteromys volans orii), this male launched itself from the top of the tree stump just fractions of a second before I pressed the shutter. The squirrel landed centimeters away from my arm on a tree that I was braced against, then rushed upward and launched itself again in pursuit of the other male.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a Pteromys volans orii flying squirrel gliding through the forest early in the morning.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a female Pteromys volans orii flying squirrel that has brought back a few small branches of an Erman's birch tree (Betula ermanii) to her nest as a takeaway meal.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel eating a male catkin from an Erman's birch tree (Betula ermanii) on a snowy day.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a male Pteromys volans orii flying squirrel sitting at the entrance to his nest.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a male Pteromys volans orii flying squirrel sitting in a favorite spot on a sunny day, looking down upon his surroundings.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel scratching his face on a branch while grooming himself early in the morning.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a male Pteromys volans orii flying squirrel attempting to enter a nest with a female inside during reproductive season. This male made repeated attempts to enter, but was stopped mid-way each time as shown here, then rejected.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a male Pteromys volans orii flying squirrel making his appearance in the late afternoon, just before heading up into the treetops to forage.
    pteromys-volans-orii-flying-squirrel...tif
  • Close-up view of a Pteromys volans orii flying squirrel finishing off a meal of male catkins from an Erman's birch tree (Betula ermanii)
    pteromys-volans-orii-flying-squirrel...tif
  • This Pteromys volans orii flying squirrel is enjoying a meal of male catkins from an Erman's birch tree (Betula ermanii).
    pteromys-volans-orii-flying-squirrel...tif
  • This is a head-on view of a Pteromys volans orii flying squirrel gliding through the forest.
    pteromys-volans-orii-flying-squirrel...tif
  • This Pteromys volans orii flying squirrel is peeking out of the entrance to its nest prior to emerging to forage.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel consuming snow for moisture after an extended period of foraging.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel leaning down to consume snow for moisture.
    pteromys-volans-orii-flying-squirrel...tif
  • Just before a Pteromys volans orii flying squirrel lands on a tree, the animals folds its wings inward and stretches its limbs toward the targeted tree. This serves to slow the animal's approach and also to prepare the animal to be able to move immediately upon landing.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel gliding through the forest canopy during light snow in winter.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel on its final approach to the nest. The squirrel has dived down from high elevation and swooped upward in an arc, bringing it to this point, which is about 1.5m from the entrance to its nest. The animal's body is flared open in order to ascend and decelerate for the final approach. Its belly is full from foraging in the treetops.
    pteromys-volans-orii-flying-squirrel...tif
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Tony Wu Underwater Photography

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