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  • A group of Lutjanus bohar twinspot snappers breaking away from the spawning aggregation to release a stream of sperm and eggs into the water. The lead fish is likely the female, with males following.
    lutjanus-bohar-snappers-spawning-pal...tif
  • Larval development of sailfin snapper (Symphorichthys spilurus) at 35 hours post-fertilization. The fertilized embryo was collected during a mass spawning aggregation involving tens of thousands of fish. Photographed in Palau.
    sailfin-snapper-embryo-symphorichthy...tif
  • Lutjanus bohar are commonly referred to as twinspot snapper, and sometimes as red snapper. As shown here, this species is capable of taking on coloration other than red. This happens during spawning aggregations, but also when fish are alone, as is the case here. The two dorsal spots can also appear and disappear. This species is an active predator, as suggested by the prominent canine teeth.
    lutjanus-bohar-twinspot-snapper-prof...tif
  • Larval development of sailfin snapper (Symphorichthys spilurus) at 10 hours post-fertilization. The fertilized embryo was collected during a mass spawning aggregation involving tens of thousands of fish. Photographed in Palau.
    sailfin-snapper-embryo-symphorichthy...tif
  • A bunch of twinspot snapper (Lutjanus bohar) rushing up to spawn, releasing a cloud of milky white sperm and eggs in the water column. Photographed in Palau.
    twinspot-snapper-spawning-lutjanus-b...tif
  • A swarm of the voracious Lutjanus bohar two-spot red snappers attacked and devoured a boxfish, the remains of which are shown here after they participated in an enormous spawning aggregation in the early morning involving thousands snappers. The attack on the boxfish was fast and furious, reminiscent of a shark feeding frenzy. I was unable to identify the species of the boxfish before it was consumed, but based on the yellow pattern, I am guessing that it was a yellow boxfish, Ostracion cubicus.
    lutanus-bohar-snapper-devouring-yell...tif
  • This is a part of a spawning aggregation comprising thousands of Bohar snapper (Lutjanus bohar) gathered in deep water in the early morning. Prior to spawning, the fish take on a range of coloration, as pictured here. Of note, the fish do not all seem to take on the same color hues or patterns. Photographed in Palau.
    lutjanus-bohar-snapper-spawning-aggr...tif
  • Twinspot snapper (Lutjanus bohar) spawning aggregation early in the morning, with a group of fish releasing a stream of gametes into the water.
    twinspot-snapper-spawning-aggregatio...tif
  • Spawning aggregation of Lutjanus bohar twinspot snapper coincidentally forming a nice heart shape, a visual metaphor for love in the ocean. Photographed in Palau.
    lutjanus-bohar-spawning-aggregation-...tif
  • Part of an enormous gathering of two-spot red snapper (Lutjanus bohar) for the purpose of mass spawning in the early morning. Thousands of fish aggregate for this spawning aggregation. Coloration of the fish seems to vary through the spawning activity, with the fish not necessarily displaying the characteristic red hue of this species. Photographed in Palau.
    two-spot-red-snapper-lutjanus-bohar-...tif
  • Single sailfin snapper, or blue-lined sea bream (Symphorichthys spilurus), swimming in the opposite direction from the rest of the school. These fish were gathered by the tens of thousands for a spawning aggregation.
    sailfin-snappers-symphorichthys-spil...tif
  • Thousands of spawning Lutjanus bohar snapper attract a number of predators, such as this blacktip shark (Carcharhinus limbatus) swimming through at rapid speed. Clouds of sperm and eggs and low light in the early morning result in low visibility, making ambush easy.
    carcharhinus-limbatus-blacktip-shark...tif
  • This is what it looks like inside a massive group of spawning Lutjanus bohar snappers. The water is milky blue from the simultaneous release of eggs and sperm by hundreds of fish in this group, which were part of an enormous gathering of thousands of fish. The visibility quickly dropped to nearly zero, with fish swimming in every direction. There were bull sharks (Carcharhinus leucas), grey reef sharks (Carcharhinus amblyrhynchos) and blackfin sharks (Carcharhinus limbatus) charging through the groups of spawning fish. Photographed in Palau.
    lutjanus-bohar-snapper-spawning-aggr...tif
  • Large school of black and white snappers (Macolor niger) patrolling just off the reef at the Blue Corner dive site in Palau
    black-and-white-snapper-Macolor-nige...tif
  • A portion of tens of thousands of blue-lined sea bream (Symphorichthys spilurus) gathered for the purpose of mass spawning at Peleliu island in Palau. These fish are normally rare in shallow water. They appear two or three times a year in large numbers for reproduction.
    blue-lined-sea-bream-aggregation-for...tif
  • After engaging in an enormous spawning aggregation in the early morning, thousands of Lutjanus bohar red snappers lingered on near a reef edge. A swarm of the snappers, comprising a couple dozen fish, attacked and devoured a boxfish, the remains of which are shown here, with the main school of snappers visible in the background. I was unable to identify the species of the boxfish before it was attacked, but based on the yellow pattern, I am guessing that it was a yellow boxfish, Ostracion cubicus.
    ostracion-cubicus-boxfish-devoured-b...tif
  • Large school of bluestripe snapper (Lutjanus kasmira) swimming along the top of a shallow reef in Palau
    bluestripe-snapper-lutjanus-kasmira-...tif
  • Massive spawning aggregation comprising thousands of two-spot red snapper (Lutjanus bohar) in the early morning. Photographed in Palau.
    lutjanus-bohar-snapper-spawning-aggr...tif
  • School of bluelined snapper (Lutjanus kasmira) at Blue Corner dive site in Palau
    bluelined-snapper-Lutjanus-kasmira-b...tif
  • Group of twinspot snappers (Lutjanus bohar) gathered as part of an early-morning spawning aggregation involving thousands of fish. Note the range of coloration displayed by individual fish, which most often appear red when seen by scuba divers.
    lutjanus-bohar-spawning-aggregation-...tif
  • This scene is typical of the chaos and pandemonium associated with mass spawning involving thousands of Lutjanus bohar red snappers. The milky clouds are massive quantities of sperm and eggs released by the frenzied fish. Photographed in Palau, in the early morning.
    twinspot-red-snapper-spawning-aggreg...tif
  • Thousands upon thousands of two-spot red snapper (Lutjanus bohar) spawning together in the early morning, producing a milky-white cloud of sperm and eggs. Photographed in Palau.
    lutjanus-bohar-snapper-mass-spawning...tif
  • Thousands of Twinspot Snapper (Lutjanus bohar) in a spawning frenzy early in the morning. Note the wide range of coloration among the individual fish. Photographed in Palau.
    lutjanus-bohar-snapper-mass-spawning...tif
  • Thousands of two-spot red snapper (Lutjanus bohar) gathered in the early morning for the purpose of spawning. Photographed in Palau.
    red-snapper-lutjanus-bohar-spawning-...tif
  • Juvenile humpback red snapper (Lutjanus gibbus). Photograph from Kokopo, Papua New Guinea.
    juvenile-humpback-red-snapper-Lutjan...tif
  • Juvenile midnight snapper against a blue background. Photograph taken in the Feni Islands, New Ireland, Papua New Guinea.
    juvenile-midnight-snapper-Macolor-ma...tif
  • Juvenile midnight snapper against a blue background. Photograph taken in the Feni Islands, New Ireland, Papua New Guinea.
    juvenile-midnight-snapper-Macolor-ni...tif
  • Profile of a blue-lined sea bream (Symphorichthys spilurus), photographed in Palau.
    blue-lined-sea-bream-symphorichthys-...tif
  • An aggregation of Lutjanus bohar snapper engaged in mass spawning. As pictured here, large groups of fish cluster together before swimming rapidly in unison to release clouds of sperm and eggs. Thousands of fish were gathered for this spawning event. Photographed in Palau.
    Lutjanus-bohar-snapper-spawning-aggr...tif
  • School of bluelined snapper (Lutjanus kasmira) at Blue Corner dive site in Palau
    bluelined-snapper-Lutjanus-kasmira-b...tif
  • Juvenile midnight snapper against a blue background. Photograph taken in the Feni Islands, New Ireland, Papua New Guinea.
    juvenile-midnight-snapper-Macolor-ma...tif
  • Twinspot snapper (Lutjanus bohar) engaged in a mass spawning aggregation early in the morning. This photograph was the winner of the Under Water category of the 52nd Wildlife Photographer of the Year competition in 2016 (#WPY52) organized by the Natural History Museum in London.
    twinspot-snapper-spawning-aggregatio...tif
  • Massive aggregation comprising tens of thousands of sailfin snapper (Symphorichthys spilurus) in preparation for group spawning. These fish normally live in deep water. They gather several times a year for this spawning phenomenon. Photographed in Palau.
    sailfin-snapper-aggregation-for-grou...tif
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