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  • This is a Japanese dwarf flying squirrel (Pteromys volans orii), a sub-species of Siberian flying squirrel. Known locally as ezo-momonga, this sub-species is found only in Hokkaido, Japan. It is primarily nocturnal. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range. This individual has just emerged shortly after sunset from its nest, a cavity in the tree pictured. After preening for a bit, the flying squirrel ascended into the canopy to forage for the night. While these squirrels are known to nest in a number of tree species, they appear to prefer Sakhalin fir trees (Abies sachalinensis).
    japanese-dwarf-flying-squirrel-at-du...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii), a sub-species of Siberian flying squirrel. Known locally as ezo-momonga, this sub-species is found only in Hokkaido, Japan. It is primarily nocturnal. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range. This individual has just emerged shortly after sunset from its nest, a cavity in the tree pictured. After preening for a bit, the flying squirrel ascended into the canopy to forage for the night. While these squirrels are known to nest in a number of tree species, they appear to prefer Sakhalin fir trees (Abies sachalinensis).
    japanese-dwarf-flying-squirrel-ptero...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This male was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii), a sub-species of Siberian flying squirrel. Known locally as ezo-momonga, this sub-species is found only in Hokkaido, Japan. It is primarily nocturnal. This indiividual has just emerged from its nest at dusk, and is taking a moment before heading out for a night of foraging. From this perspective, it is possible to see how the squirrel grasps and maintains a grip on delicate branches. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range.
    japanese-dwarf-flying-squirrel-at-du...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii), a sub-species of Siberian flying squirrel. that has just emerged from its nest before setting out for a night of foraging. Known locally as ezo-momonga, this sub-species is found only in Hokkaido, Japan. It is primarily nocturnal. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range.
    japanese-dwarf-flying-squirrel-at-du...tif
  • This Japanese dwarf flying squirrel (Pteromys volans orii) is just emerging from its nest in the trunk of a tree at dusk to prepare for a night of foraging. Found only in Hokkaido, Japan, these animals prefer small holes to their nests, as this provides a measure of protection from predators like owls and martens. This particular nest entrance is also partially covered by a large white fungus, which obscures it from view. This animal is a sub-species of Siberian flying squirrel and is known locally as ezo-momonga.
    japanese-dwarf-flying-squirrel-emerg...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii) that has just come out from its nest., entrance visible to left and above the animal’s head. These flying squirrels are nocturnal, but occasionally emerge during daylight hours.
    japanese-dwarf-flying-squirrel-20200...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii) peeking out from its nest in the trunk of a tree. Checking the surroundings in this manner prior to emerging at dusk for a night of foraging is common behaviour. Found only in Hokkaido, Japan, these animals prefer small holes to their nests, as this provides a measure of protection from predators like owls and martens. This particular nest entrance is also partially covered by a large white fungus, which obscures it from view. This animal is a sub-species of Siberian flying squirrel and is known locally as ezo-momonga.
    japanese-dwarf-flying-squirrel-at-du...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii) sitting on a branch. This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii), a sub-species of Siberian flying squirrel, emerging from its nest at dusk to set out for a night of foraging. Known locally as ezo-momonga, this sub-species is found only in Hokkaido, Japan. It is primarily nocturnal. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range.
    japanese-dwarf-flying-squirrel-at-du...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii), a sub-species of Siberian flying squirrel. getting ready to leave its nest to set out for a night of foraging. Known locally as ezo-momonga, this sub-species is found only in Hokkaido, Japan. It is primarily nocturnal. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range.
    japanese-dwarf-flying-squirrel-at-du...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii), a sub-species of Siberian flying squirrel, peeking out from its nest before emerging for a night of foraging. Known locally as ezo-momonga, this sub-species is found only in Hokkaido, Japan. It is primarily nocturnal. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range.
    japanese-dwarf-flying-squirrel-at-du...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii) taking a moment to gather itself after emerging from its nest to head out for a night of foraging. Known locally as ezo-momonga, this sub-species of Siberian flying squirrel is found only in Hokkaido, Japan. It is primarily nocturnal. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range.
    japanese-dwarf-flying-squirrel-at-du...tif
  • During the breeding season, male Japanese dwarf flying squirrels (Pteromys volans orii) sometimes confront one another, usually in the context of competition for a female. Here, the one above has just snuck up behind the individual below, which made a quick turn to look up at the intruder. The animal looking up was in the midst of enjoying a snack, the flowers of a Japanese alder (Alnus japonica) that are visible in his mouth. A frantic battle between the two males soon ensued.
    japanese-dwarf-flying-squirrel-males...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii) that has found a comfortable place to perch on an overcast day after emerging from its nest. These animals line the inside of their nests with soft materials for comfort and warmth. Some of this nest lining is visible on this squirrel’s head.
    japanese-dwarf-flying-squirrel-20200...tif
  • Japanese dwarf flying squirrel (Pteromys volans orii) perched at the juncture of a small branch and tree trunk. This species is normally nocturnal. This one appeared during briefly during the day, but soon returned to its nest.
    japanese-dwarf-flying-squirrel-20200...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii) that has just emerged from its nest at dusk to set out for a night of foraging.
    japanese-dwarf-flying-squirrel-20200...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii) that has found a comfortable place to perch, just as mid-afternoon sunlight broke through a cloud-filled sky. These animals are nocturnal, but occasionally emerge from their nests during daylight hours.
    japanese-dwarf-flying-squirrel-20200...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii) shaking itself off at the entrance of its nest.
    japanese-dwarf-flying-squirrel-20200...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii) grooming itself while resting in a tree in the golden light of late afternoon. Although this species is primarily nocturnal, individuals occasionally emerge from their nests during daylight hours. This individual was well hidden in a shaded area covered by a tangle of small branches.
    japanese-dwarf-flying-squirrel-groom...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii) that has found a comfortable place to perch on an overcast day.
    japanese-dwarf-flying-squirrel-20200...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii) resting in a tree in the golden light of late afternoon. Although this species is primarily nocturnal, individuals occasionally emerge from their nests during daylight hours. This individual was well hidden in a shaded area covered by a tangle of small branches.
    japanese-dwarf-flying-squirrel-20200...tif
  • When a Japanese dwarf flying squirrel (Pteromys volans orii) emerges from its nest at dusk, one of the first orders of business is often to relieve itself. The individual pictured here is defecating (pellets visible in the appropriate location) while grasping a small twig for stability. Flying squirrel droppings are used in traditional Chinese herbal medicine for ailments related to blood circulation and abdominal pain. The droppings are known as Wu Ling Zhi.
    japanese-dwarf-flying-squirrel-defec...tif
  • This Japanese dwarf flying squirrel (Pteromys volans orii) snacking on a Japanese alder (Alnus japonica) flower, one of this species’ preferred foods.
    japanese-dwarf-flying-squirrel-20200...tif
  • In the midst of competition for the right to mate with a female Japanese dwarf flying squirrel (Pteromys volans orii), this male launched itself from the top of the tree stump just fractions of a second before I pressed the shutter. The squirrel landed centimeters away from my arm on a tree that I was braced against, then rushed upward and launched itself again in pursuit of the other male.
    japanese-dwarf-flying-squirrel-male-...tif
  • A female Japanese dwarf flying squirrel (Pteromys volans orii) in oestrus often attracts the attention of more than one male. When that happens, males chase one another, leap from and glide among trees, and sometimes engage in physical skirmishes. The male in the background of this image is stalking the male in the foreground in an attempted sneak-attack during one such contest for an eligible female.
    japanese-dwarf-flying-squirrel-males...tif
  • This is the moment that one male Japanese dwarf flying squirrel (Pteromys volans orii) ambushed a rival male, sneaking up from behind and leaping on on top of the other male. Male rivalry and competition like this takes place during the reproductive season. The action is rapid, often spanning wide areas as the animals leap and glide among trees.
    japanese-dwarf-flying-squirrel-males...tif
  • This is a pair of Japanese dwarf flying squirrels (Pteromys volans orii) engaged in copulation. The crack in the middle of the tree is the entrance to their nest.
    japanese-dwarf-flying-squirrel-copul...tif
  • This is a male and female pair of Japanese dwarf flying squirrels (Pteromys volans orii), with one inside the nest and the other above. This female (inside the nest) is in oestrus. The male is standing guard. Another male approached repeatedly, resulting in heated confrontation, with the males chasing one another, occasionally leaping and gliding among trees.
    japanese-dwarf-flying-squirrel-matin...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-20200...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous. The squirrel’s paddle-like tail is visible from this angle, flopped over its back. This animal uses its tail somewhat like a rudder during flight, helping to adjust direction.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This male was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous. The squirrel paused for a moment to groom himself.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This male was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This male was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This male was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous. The squirrel took a moment here to scratch his back.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-20200...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This male was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii), gliding from the top of one tree to another. Known locally as ezo-momonga, this sub-species of Siberian flying squirrel is found only in Hokkaido, Japan. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range. Although these animals are primarily nocturnal, they sometimes emerge during the day. Usually this is for brief durations to urinate or defecate. In this instance, this particular individual was one of two males competing for the right to mate with a female. Such reproduction-related competition sometimes takes place during daylight hours.
    japanese-dwarf-flying-squirrel-airbo...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii), looking out from his nest. This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This male was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • Side view of a Japanese dwarf flying squirrel (Pteromys volans orii), which is primarily a nocturnal species, but males like this sometimes emerge during daylight hours of the reproductive season.
    japanese-dwarf-flying-squirrel-male-...tif
  • Pictured here is a pair of a Japanese dwarf flying squirrel (Pteromys volans orii), one peeking out from the nest, and another perched on a branch above. These squirrels are preparing for a night of foraging, having emerged shortly after sunset. Known locally as ezo-momonga, this sub-species is a  sub-species of Siberian flying squirrel and is found only in Hokkaido, Japan. It is primarily nocturnal. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range.
    japanese-dwarf-flying-squirrels-at-d...tif
  • This series of six images shows a pair of Japanese dwarf flying squirrels (Pteromys volans orii) that have just emerged from a nest in the tree. The male pursues the female; the pair copulate; and the female returns to the nest. Over the course of a day or so, the male copulates as many times as possible. Rival males sometimes approach, which can result in a heated battle between males.
    japanese-dwarf-flying-squirrels-copu...tif
  • This is a pair of Japanese dwarf flying squirrels (Pteromys volans orii) engaged in copulation. This is an extended process spanning many hours, during which the male often needs to fend off rivals while engaging in repeated bouts of copulation.
    japanese-dwarf-flying-squirrels-mati...tif
  • This is a pair of Japanese dwarf flying squirrels (Pteromys volans orii) engaged in copulation. This is an extended process spanning many hours, during which the male often needs to fend off rivals while engaging in repeated bouts of copulation.
    japanese-dwarf-flying-squirrels-mati...tif
  • This is a pair of Japanese dwarf flying squirrels (Pteromys volans orii) engaged in copulation. This is an extended process spanning many hours, during which the male often needs to fend off rivals while engaging in repeated bouts of copulation.
    japanese-dwarf-flying-squirrels-mati...tif
  • This is a pair of Japanese dwarf flying squirrels (Pteromys volans orii) engaged in copulation. This is an extended process spanning many hours, during which the male often needs to fend off rivals while engaging in repeated bouts of copulation. As shown here, the female can, and often does, put up some measure of resistance, occasionally dislodging the male completely. The male’s penis is visible.
    japanese-dwarf-flying-squirrels-mati...tif
  • This is a pair of Japanese dwarf flying squirrels (Pteromys volans orii) engaged in copulation. This is an extended process spanning many hours, during which the male often needs to fend off rivals while engaging in repeated bouts of copulation.
    japanese-dwarf-flying-squirrels-mati...tif
  • This is a pair of Japanese dwarf flying squirrels (Pteromys volans orii) engaged in copulation. This is an extended process spanning many hours, during which the male often needs to fend off rivals while engaging in repeated bouts of copulation. As shown here, the female can, and often does, put up some measure of resistance, occasionally dislodging the male completely. The male’s penis is visible.
    japanese-dwarf-flying-squirrels-mati...tif
  • This is a pair of Japanese dwarf flying squirrels (Pteromys volans orii) engaged in copulation. This is an extended process spanning many hours, during which the male often needs to fend off rivals while engaging in repeated bouts of copulation.
    japanese-dwarf-flying-squirrels-mati...tif
  • This is a pair of Japanese dwarf flying squirrels (Pteromys volans orii), photographed during breeding season. The individual higher up on the tree is a female in oestrus. In the first two images in this sequence of three, the male approaches the female from behind to attempt copulation. In the third image, the female spins around to indicate in no uncertain terms that she is not ready. The pair’s nest is in the crack in the tree.
    japanese-dwarf-flying-squirrels-copu...tif
  • This is a Pteromys volans orii Japanese dwarf flying squirrel foraging at dawn. It is high in the canopy, around 15-20m, not far from its nest. The tree is a Japanese elm (Ulmus davidiana var. japonica), one of several prime food sources for this species. As seen here, these flying squirrels consume the leaf buds of this tree.
    japanese-flying-squirrel-pteromys-vo...tif
  • This Japanese flying squirrel (Pteromys volans orii) is perched on a comfortable, safe location high above the ground, where it is eating one of its favorite foods, the buds of a Japanese elm tree (Ulmus davidiana var. japonica). Normally nocturnal, these squirrels are sometimes active in the early hours of daylight.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii) flying through the forest on a cold winter day. Usually nocturnal, these animals sometimes become active in the day during reproductive season.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a Pteromys volans orii Japanese dwarf flying squirrel flying through the forest during a flurry of activity involving five individuals, two females and three males. The two females were in estrus.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a Pteromys volans orii Japanese dwarf flying squirrel gliding across the top of the tree canopy after sunset. The moon is visible, six days prior to being full. This individual was part of a ruckus of reproductive season activity stretching over many hours that involved two females and three males.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a Pteromys volans orii Japanese dwarf flying squirrel male perched at the top entrance to a nest. On this day, he was guarding a female inside the nest. She was in estrus, and the male was warding off other males. This pair mated several times throughout the day, but the female also mated with another male while this one was unaware. This photo is first in a series of three which illustrate the height of the nest.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a Pteromys volans orii Japanese dwarf flying squirrel male perched at the top entrance to a nest. On this day, he was guarding a female inside the nest. She was in estrus, and the male was warding off other males. This pair mated several times throughout the day, but the female also mated with another male while this one was unaware. This photo is one in a series of three which illustrate the height of the nest.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a Pteromys volans orii Japanese dwarf flying squirrel male perched at the top entrance to a nest. On this day, he was guarding a female inside the nest. She was in estrus, and the male was warding off other males. This pair mated several times throughout the day, but the female also mated with another male while this one was unaware. This photo is third in a series of three which illustrate the height of the nest.
    japanese-flying-squirrel-pteromys-vo...tif
  • Pictured here is a pair of Japanese dwarf flying squirrels (Pteromys volans orii) shortly before mating. The female is in front. She has lifted her tail, letting the male check her status by smell. They mated one hour after this photo was taken. This perch is stained with urine, and there are feces visible. This is an indication that it is frequented by these and possibly other flying squirrels.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a pair of Japanese dwarf flying squirrels (Pteromys volans orii). The one in front is male. The one peeking out from behind is female. They are situated at the top entrance to their nest, caressed by warm light from the setting winter sun. This pair mated several times throughout the day. There was another male in the area, a challenger. The female mated with the other male as well, when this one was not looking.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a Pteromys volans orii flying squirrel enjoying a meal of Japanese elm buds (Ulmus davidiana var. japonica) while perched more than 20m above the ground. Note the squirrel's perfect balance while seated at the end of tiny branches.
    pteromys-volans-orii-flying-squirrel...tif
  • This Pteromys volans orii flying squirrel is relishing fresh buds of a small branch from a Japanese elm tree (Ulmus davidiana var. japonica).
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel carrying a small branch of Japanese elm (Ulmus davidiana var. japonica) back to its nest. This individual had spent over an hour in the early morning at the top of 20m tall elm trees feasting on fresh buds. This branch is a take-away meal for enjoyment back home.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel returning to its nest at dawn after foraging high in the canopy during the night. It is carrying a small branch with leaf buds of a Japanese elm (Ulmus davidiana var. japonica), one of several prime food sources for this species.
    japanese-flying-squirrel-pteromys-vo...tif
  • This Pteromys volans orii flying squirrel has carried a few sticks of Japanese elm (Ulmus davidiana var. japonica) back to its nest at the end of foraging to enjoy a takeaway meal before going inside.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel en route back to its nest after foraging at the top of 20m+ tall Japanese elm trees (Ulmus davidiana var. japonica) for an extended period of time during snowfall. The squirrel is carrying a branch with fresh elm buds back to its nest—a takeaway meal to relish at home.
    pteromys-volans-orii-flying-squirrel...tif
  • This Japanese flying squirrel (Pteromys volans orii) has just emerged from its nest at dusk, before setting out for a night of a foraging in the tree tops.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel perched at the top of a 20+m Japanese elm tree (Ulmus davidiana var. japonica) early in the morning. Fresh buds of this elm species are one of the preferred foods of these squirrels.
    pteromys-volans-orii-flying-squirrel...tif
  • This Japanese flying squirrel (Pteromys volans orii) has just emerged from its nest during daylight hours. Although these animals are nocturnal, they sometimes come out during the day.
    pteromys-volans-orii-flying-squirrel...tif
  • The Pteromys volans orii flying squirrel inside the nest has brought a few sticks of Japanese elm (Ulmus davidiana var. japonica) back at the end of foraging to enjoy before going inside. Sensing opportunity, the other occupant of the nest has also returned, hoping to share in the bounty. A squabble ensued.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel eating one of its favorite foods—catkins from a Japanese alder tree (Alnus japonica). This individual had descended to ground level during a winter evening to forage for fallen catkins. Descending in this manner is unusual behavior.
    pteromys-volans-orii-flying-squirrel...tif
  • Before emerging from their nests, Pteromys volans orii flying squirrels often do a visual check of their surroundings. In this instance, the entrance to the nest was so small that the squirrel's face appeared larger than the hole, resulting in an amusing peekaboo effect.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a Pteromys volans orii flying squirrel male grooming itself after emerging from his nest on an early winter morning. This male shared the nest with a female. The pair mated 10 days after this photo was taken.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a Pteromys volans orii flying squirrel male grooming itself after emerging from his nest on an early winter morning. This male shared the nest with a female. The pair mated 10 days after this photo was taken.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a Pteromys volans orii flying squirrel male grooming itself after emerging from his nest on an early winter morning. This male shared the nest with a female. The pair mated 10 days after this photo was taken.
    japanese-flying-squirrel-pteromys-vo...tif
  • During reproductive season, Pteromys volans orii flying squirrel males sometimes compete for females that have gone into estrus. Pictured here are two male rivals. One is guarding the entrance to a hole in a tree. Inside is the female. The other male is sneaking up from the other side of the tree with the intention of challenging the ensconced male.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a pair of Pteromys volans orii flying squirrels that have just emerged from their nest early in the morning during the winter reproductive season. The individual on the left is female, right male. After preening together, the pair ascended into the canopy to forage. They mated 10 days after this photo was taken.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a Pteromys volans orii flying squirrel peeking out from its den in a Sakhalin fir tree (Abies sachalinensis) in the soft light of the early winter morning.
    japanese-flying-squirrel-pteromys-vo...tif
  • Pictured here are two male Pteromys volans orii flying squirrels. The one inside the tree has just peeked out from the nest which he has occupied with a female that has gone into estrus. Caught sneaking up to try to enter the nest to compete for the female, the male on the tree has just jumped away to avoid a confrontation.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a Pteromys volans orii flying squirrel male pausing after grooming himself during an early winter morning. This male shared the nest with a female, which joined him shortly after this photo was taken. The pair mated 10 days after this photo was taken.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a male Pteromys volans orii flying squirrel standing guard at the entrance of a nest. Inside is a female that has entered estrus. This has attracted the attention of another male. At this point, the interloper male had made numerous attempts to mate with the female, hence the protective position of this male. The two males faced off a number of times. Interlopers are sometimes successful in mating with females while the primary male is unaware or otherwise occupied.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a Pteromys volans orii flying squirrel male grooming itself after emerging from his nest on an early winter morning. This male shared the nest with a female. The pair mated 10 days after this photo was taken.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a portrait of a Pteromys volans orii flying squirrel male illuminated by the light of an early winter morning. He had just emerged from his nest, followed shortly thereafter by a female. The pair mated 10 days after this photo was taken.
    japanese-flying-squirrel-pteromys-vo...tif
  • Competition between rival males for the right to mate with a female flying squirrel (Pteromys volans orii) in estrus can become heated. The two males pictured here chased one another up and down the tree, culminating in one executing a full body-slam punch, sending the other into the air.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a pair of Pteromys volans orii flying squirrels that have just emerged from their nest early in the morning during the winter reproductive season. The individual on the left is female, right male. After preening together, the pair ascended into the canopy to forage. They mated 10 days after this photo was taken.
    japanese-flying-squirrel-pteromys-vo...tif
  • This male and female Pteromys volans orii flying squirrels have paired up for the reproductive season. They had just finished foraging for food high in the canopy and were sitting together like this during light snow. The pair mated a few days after this photograph was taken. The male is on the right, female left.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a male Pteromys volans orii flying squirrel gliding through the canopy in mid-afternoon. This species is primarily nocturnal. During the peak of the reproductive season, they can become active during the day when there is competition among males for a female in estrus, as was the case here.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a pair of Pteromys volans orii flying squirrels that have just emerged from their nest early in the morning during the winter reproductive season. The individual on the left is female, right male. After preening together, the pair ascended into the canopy to forage. They mated 10 days after this photo was taken.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a pair of Pteromys volans orii flying squirrels that have just emerged from their nest early in the morning during the winter reproductive season. The individual facing the camera is male. After preening together, the pair ascended into the canopy to forage. They mated 10 days after this photo was taken.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a male Pteromys volans orii flying squirrel perched in a safe location high up in the canopy. Behind this knot in the tree was the nest. The squirrel was positioned at the top entrance to the nest, which gave him the ideal vantage point from which to survey the surroundings and keep a lookout for two rival males that were attempting to get the attention of the female inside. Many battles ensued, with much chasing and lots of flying.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel flying through trees.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel flying directly overhead.
    pteromys-volans-orii-flying-squirrel...tif
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