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  • Juvenile Sika Deer (Cervus nippon yesoensis) foraging for food in the winter. Here the deer is nibbling on Japanese rose (Rosa rugosa). This is the Hokkaido subspecies of Cervus nippon. Photographed in Biei, Hokkaido, Japan. エゾシカ, 美瑛、北海道
    japanese-sika-deer-hokkaido-subspeci...tif
  • This species of crayfish, known as zarigani (Cambaroides japonicus) is endemic to Japan, living in Hokkaido and northern Honshu. They live in areas of low temperature with high water purity. This individual was missing its right front claw.
    zarigani-crayfish-cambaroides-japoni...tif
  • Shirahige no Taki Waterfall (白ひげの滝) in winter. Photographed in Biei, Hokkaido, Japan.
    waterfall-biei-hokkaido-japan-shirah...tif
  • Japanese larch trees (Larix kaempferi) in the blue ice of Blue Pond in Biei, Hokkaido, Japan frozen over during winter.
    japanese-larch-trees-frozen-blue-pon...tif
  • Waterfall named Fudou no Taki (不動の滝) after the first major snowfall of the winter. Photographed in Biei, Hokkaido, Japan.
    waterfall-in-winter-biei-hokkaido-ja...tif
  • Blue Pond in Biei, Hokkaido, Japan in the winter, with snow falling. The pond is manmade, and the trees growing through the water are Japanese larch (Larix kaempferi), deciduous conifers. 北海道の美瑛にある青い池です。
    blue-pond-in-winter-with-snow-hokkai...tif
  • Blue Pond in Biei, Hokkaido, Japan in the winter, with snow falling. The pond is manmade, and the trees growing through the water are Japanese larch (Larix kaempferi), deciduous conifers. 北海道の美瑛にある青い池です。
    blue-pond-in-winter-biei-hokkaido-ja...tif
  • Aerial panorama of autumn foliage in southern Hokkaido, Japan
    hokkaido-japan-autumn-foliage-aerial...tif
  • Icicles in the early morning in Hokkaido, Japan
    icicles-pre-dawn-winter-hokkaido-jap...tif
  • This is a waterfall in Biei, Hokkaido, Japan called Shirahige no Taki (白ひげの滝), photographed in winter after the first snowfall of the season.
    waterfall-biei-hokkaido-shirahige-no...tif
  • Steller's sea eagle (Haliaeetus pelagicus) looking out to sea from a comfortable perch on a tree branch. Photographed in Rausu, Hokkaido, Japan.
    stellers-sea-eagle-haliaeetus-pelagi...tif
  • Red fox, Sakhalin subspecies (Vulpes vulpes schrencki) foraging for food in the winter snow of Hokkaido. Photographed in Shiretoko National Park.
    red-fox-sakhalin-subspecies-vulpes-v...tif
  • Female Japanese sika deer (Cervus nippon yesoensis) with young fawn. The deer were foraging for food during the winter in Utoro, Hokkaido, Japan.
    japanese-sika-deer-Cervus-nippon-yes...tif
  • Blakiston’s Fish Owl (Bubo blackistoni) at night, with a small masu salmon (Oncorhynchus masou masou), also known as yamame, in its mouth. Photographed in Rausu, Hokkaido, Japan. This owl is accustomed to people and is fed daily.
    endangered-blakistons-fish-owl-bubo-...tif
  • Blakiston’s Fish Owl (Bubo blackistoni) at night. Photographed in Rausu, Hokkaido, Japan. This owl is accustomed to people and is fed daily.
    endangered-blakistons-fish-owl-bubo-...tif
  • Red fox, Sakhalin subspecies (Vulpes vulpes schrencki) foraging for food in the winter snow of Hokkaido. Photographed in Shiretoko National Park.
    red-fox-sakhalin-subspecies-vulpes-v...tif
  • This is an aerial panorama showing some of the sulfur and steam vents at Mount Io-zan in Hokkaido, Japan. The volcano rises 512m within the Kussharo caldera. Lake Kussharo is just visible in the far right background.
    sulfur-steam-vent-volcano-mount-ioza...tif
  • This is an aerial photo showing some of the sulfur and steam vents at Mount Io-zan in Hokkaido, Japan. The volcano rises 512m within the Kussharo caldera.
    sulfur-steam-vent-volcano-mount-ioza...tif
  • This is an aerial photo showing some of the sulfur and steam vents at Mount Io-zan in Hokkaido, Japan. The volcano rises 512m within the Kussharo caldera.
    sulfur-steam-vent-volcano-mount-ioza...tif
  • Pictured here are two pairs of Japanese cranes (Grus japonensis)  in Hokkaido, Japan crossing paths while engaged in pair-bonding. This species is found in Siberia, Northeast China, Mongolia, Korea and northern Japan. The population in northern Japan is mostly non-migratory, remaining resident on the island of Hokkaido throughout the year. This species is listed as Endangered on the IUCN Red List.
    grus-japonensis-japanese-red-crowned...tif
  • This is a pair of mature Japanese cranes (Grus japonensis) engaged in a pair-bonding performance in Hokkaido, Japan. This species is found in Siberia, Northeast China, Mongolia, Korea and northern Japan. The population in northern Japan is mostly non-migratory, remaining resident on the island of Hokkaido throughout the year. This species is listed as Endangered on the IUCN Red List.
    grus-japonensis-japanese-red-crowned...tif
  • This is a pair of mature Japanese cranes (Grus japonensis) engaged in a pair-bonding performance in Hokkaido, Japan. This species is found in Siberia, Northeast China, Mongolia, Korea and northern Japan. The population in northern Japan is mostly non-migratory, remaining resident on the island of Hokkaido throughout the year. This species is listed as Endangered on the IUCN Red List.
    grus-japonensis-japanese-red-crowned...tif
  • This is a pair of mature Japanese cranes (Grus japonensis) engaged in a pair-bonding display during a snowstorm in Hokkaido, Japan. This species is found in Siberia, Northeast China, Mongolia, Korea and northern Japan. The population in northern Japan is mostly non-migratory, remaining resident on the island of Hokkaido throughout the year. This species is listed as Endangered on the IUCN Red List.
    grus-japonensis-pair-bonding-snowsto...tif
  • Blakiston’s Fish Owl (Bubo blackistoni) at night, with a small masu salmon (Oncorhynchus masou masou), also known as yamame, in its mouth. Photographed in Rausu, Hokkaido, Japan. This owl is wild, but is habituated to people and is fed daily.
    endangered-blakistons-fish-owl-bubo-...tif
  • This is a Strix uralensis japonica owl in its nest, feathers being ruffled by a strong breeze.
    strix-uralensis-japonica-ural-owl-ez...tif
  • Portrait of a female Japanese spotted deer (Cervus nippon yesoensis), photographed in the snow at Shiretoko National Park, Utoro, Hokkaido, Japan.
    japanese-spotted-deer-portrait-cervu...tif
  • Japanese spotted deer (Cervus nippon yesoensis) foraging for food during winter, along the edge of a slope overlooking the Okhotsk Sea. Photographed in Shiretoko National Park, Utoro, Hokkaido, Japan.
    japanese-spotted-deer-portrait-cervu...tif
  • Japanese spotted deer (Cervus nippon yesoensis) with young fawn and two more deer in the background. The deer were foraging for food during winter. Photographed in Shiretoko National Park, Utoro, Hokkaido, Japan.
    japanese-spotted-deer-cervus-nippon-...tif
  • This is a pair of Ural owls (Strix uralensis japonica) from the population found in Hokkaido, Japan. The male on the left is grooming the female. The female will soon lay eggs and raise chicks in the coming spring.
    strix-uralensis-japonica-ural-owl-ez...tif
  • This is a female Lottia emydia limpet sending a stream of eggs into the water during spawning. These sea snails aggregate in order to spawn en masse. Photographed in Hokkaido, Japan.
    lottia-emydia-limpet-spawning-japan-...tif
  • This is a pair of Eurasian Coots (Fulica atra) grazing for seagrass in a shallow inlet in Hokkaido, Japan.
    fulica-atra-coots-eating-seagrass-ja...tif
  • This is a Chirolophis saitone, called akiginpo in Japanese. This fish is a species of prickleback. It reaches a size of about 10cm, and it lives in shallow coastal waters in the Sea of Japan and around Hokkaido.
    chirolophis-saitone-profile-japan-20...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous. The squirrel took a moment here to scratch his back.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-20200...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii) sitting on a branch. This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous. The squirrel’s paddle-like tail is visible from this angle, flopped over its back. This animal uses its tail somewhat like a rudder during flight, helping to adjust direction.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii), looking out from his nest. This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This male was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii), gliding from the top of one tree to another. Known locally as ezo-momonga, this sub-species of Siberian flying squirrel is found only in Hokkaido, Japan. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range. Although these animals are primarily nocturnal, they sometimes emerge during the day. Usually this is for brief durations to urinate or defecate. In this instance, this particular individual was one of two males competing for the right to mate with a female. Such reproduction-related competition sometimes takes place during daylight hours.
    japanese-dwarf-flying-squirrel-airbo...tif
  • This is a female Lottia emydia limpet sending a stream of eggs into the water during spawning. These sea snails aggregate in order to spawn en masse. Photographed in Hokkaido, Japan.
    lottia-emydia-limpet-spawning-japan-...tif
  • This is a sulfurous steam vent at Mount Io-zan in eastern Hokkaido, Japan. Mount Io-zan is a 512-meter high volcano sitting inside the Kussharo caldera.
    sulfur-steam-vent-volcano-mount-ioza...tif
  • Whooper swan (Cygnus cygnus) visiting wintering grounds in Hokkaido, Japan. The swan is turned tail-end up while foraging for food.
    cygnus-cygnus-whooper-swan-foraging-...tif
  • Pictured in the foreground is a male chum salmon (Oncorhynchus keta) on his last legs during the spawning run. His body has deteriorated considerably. His eye is opaque. Yet, he was still vying to spawn with females. A more intact male is visible in the background. Photographed in Hokkaido, Japan.
    oncorhynchus-keta-chum-salmon-spawni...tif
  • These are chum salmon (Oncorhynchus keta) swimming in a shallow river in Hokkaido, Japan. The fish are preparing to spawn after spending one to three years at sea. The individual in the foreground is female, the larger two in the background males. Females dig shallow nests at the edges of waterflow to deposit eggs, which are fertillized by a partner male, sometimes multiple males. The female covers fertilized eggs with gravel.
    oncorhynchus-keta-chum-salmon-spawni...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This male was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous. The squirrel paused for a moment to groom himself.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This male was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii), a sub-species of Siberian flying squirrel. Known locally as ezo-momonga, this sub-species is found only in Hokkaido, Japan. It is primarily nocturnal. This indiividual has just emerged from its nest at dusk, and is taking a moment before heading out for a night of foraging. From this perspective, it is possible to see how the squirrel grasps and maintains a grip on delicate branches. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range.
    japanese-dwarf-flying-squirrel-at-du...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii), a sub-species of Siberian flying squirrel. that has just emerged from its nest before setting out for a night of foraging. Known locally as ezo-momonga, this sub-species is found only in Hokkaido, Japan. It is primarily nocturnal. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range.
    japanese-dwarf-flying-squirrel-at-du...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii), a sub-species of Siberian flying squirrel. getting ready to leave its nest to set out for a night of foraging. Known locally as ezo-momonga, this sub-species is found only in Hokkaido, Japan. It is primarily nocturnal. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range.
    japanese-dwarf-flying-squirrel-at-du...tif
  • This Japanese dwarf flying squirrel (Pteromys volans orii) is just emerging from its nest in the trunk of a tree at dusk to prepare for a night of foraging. Found only in Hokkaido, Japan, these animals prefer small holes to their nests, as this provides a measure of protection from predators like owls and martens. This particular nest entrance is also partially covered by a large white fungus, which obscures it from view. This animal is a sub-species of Siberian flying squirrel and is known locally as ezo-momonga.
    japanese-dwarf-flying-squirrel-emerg...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii), a sub-species of Siberian flying squirrel. Known locally as ezo-momonga, this sub-species is found only in Hokkaido, Japan. It is primarily nocturnal. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range. This individual has just emerged shortly after sunset from its nest, a cavity in the tree pictured. After preening for a bit, the flying squirrel ascended into the canopy to forage for the night. While these squirrels are known to nest in a number of tree species, they appear to prefer Sakhalin fir trees (Abies sachalinensis).
    japanese-dwarf-flying-squirrel-ptero...tif
  • This is a species of flying squirrel called Ezo Momonga in Japanese (Pteromys volans orii). It is a sub-species of Siberian flying squirrels that is found on the island of Hokkaido in Japan. These squirrels are primarily nocturnal and usually shy, though the squirrels can occasionally be seen during the day. They prefer to take up residence in holes made by woodpeckers. This individual has just emerged from its hole in a Sakhalin fir tree  (Abies sachalinensis) at dusk, preparing for a night of foraging. Their diet primarily comprises leaves, seeds, cones, buds, sprouts, nuts and berries. This species does not hibernate.
    ezo-momonga-pteromys-volans-orii-fly...tif
  • This is a species of flying squirrel called Ezo Momonga in Japanese (Pteromys volans orii). It is a sub-species of Siberian flying squirrels that is found on the island of Hokkaido in Japan. These squirrels are primarily nocturnal and usually shy, though the squirrels can occasionally be seen during the day. They prefer to take up residence in holes made by woodpeckers, as seen here. This individual is just emerging from a hole in a Sakhalin fir tree (Abies sachalinensis) at dusk, in preparation for a night of foraging. These small animals are particularly wary when they emerge, as they must be on the lookout for predators like owls and weasels. Their diet primarily comprises leaves, seeds, cones, buds, sprouts, nuts and berries. This species does not hibernate.
    ezo-momonga-pteromys-volans-orii-fly...tif
  • This is a species of flying squirrel called Ezo Momonga in Japanese (Pteromys volans orii). It is a sub-species of Siberian flying squirrels that is found on the island of Hokkaido in Japan. These squirrels are primarily nocturnal and usually shy, though the squirrels can occasionally be seen during the day. They prefer to take up residence in holes made by woodpeckers. This individual has just emerged from its hole in a Sakhalin fir tree  (Abies sachalinensis) at dusk, preparing for a night of foraging. Their diet primarily comprises leaves, seeds, cones, buds, sprouts, nuts and berries. This species does not hibernate.
    ezo-momonga-pteromys-volans-orii-fly...tif
  • This is a juvenile Japanese crane (Grus japonensis) in a snowstorm. It is holding a rock in its beak, one that it picked up while foraging for food with its parents. This species is found in Siberia, Northeast China, Mongolia, Korea and northern Japan. The population in northern Japan is mostly non-migratory, remaining resident on the island of Hokkaido throughout the year. This species is listed as Endangered on the IUCN Red List.
    grus-japonensis-japanese-red-crowned...tif
  • This is a species of flying squirrel called Ezo Momonga in Japanese (Pteromys volans orii). It is a sub-species of Siberian flying squirrels that is found on the island of Hokkaido in Japan. These squirrels are primarily nocturnal and usually shy, though the squirrels can occasionally be seen during the day. They prefer to take up residence in holes made by woodpeckers. Their diet primarily comprises leaves, seeds, cones, buds, sprouts, nuts and berries. This individual is nibbling on the soft parts of a Sakhalin fir tree (Abies sachalinensis), known as todomatsu in Japanese. This species does not hibernate.
    ezo-momonga-pteromys-volans-orii-fly...tif
  • This is a cluster of Lottia emydia limpets gathered for group spawning. One is just starting to lift itself up from the rock to spawn. Above these limpets is a cloud of eggs released by nearby female limpets. Photographed in Hokkaido, Japan.
    lottia-emydia-limpet-spawning-japan-...tif
  • This is a female Lottia emydia limpet sending a stream of eggs into the water during spawning. These sea snails aggregate in order to spawn en masse. Photographed in Hokkaido, Japan.
    lottia-emydia-limpet-spawning-japan-...tif
  • This is a whooper swan (Cygnus cygnus) in flight. Many of these birds spend their winters in Hokkaido, Japan, then fly north again with the arrival of warmer weather in spring.
    cygnus-cygnus-whooper-swan-japan-202...tif
  • Pictured here is a pair of Ural owls (Strix uralensis japonica), known locally in Hokkaido as Ezo fukurou. The smaller individual on the lower left is the male. This pair was preparing to have chicks.
    strix-uralensis-japonica-ural-owl-ez...tif
  • This is a flock of 30 whooper swans (Cygnus cygnus) heading north from Hokkaido, commencing their migration to higher latitudes for the warmer months of the year.
    cygnus-cygnus-whooper-swan-migration...tif
  • This is a long-tailed tit (Aegithalos caudatus) with winter plumage. Photographed in Hokkaido, Japan.
    long-tailed-tit-aegithalos-caudatus-...tif
  • This is a long-tailed tit (Aegithalos caudatus) with winter plumage. Photographed in Hokkaido, Japan.
    long-tailed-tit-aegithalos-caudatus-...tif
  • Whooper swan (Cygnus cygnus) in Hokkaido wintering grounds eating seagrass that it grabbed while wading in a shallow bay
    cygnus-cygnus-whooper-swan-japan-202...tif
  • This is a male chum salmon (Oncorhynchus keta). Despite his poor condition, the male was still swimming and competing with other males to spawn with females. Photographed in Hokkaido, Japan.
    oncorhynchus-keta-chum-salmon-spawni...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-20200...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii) peeking out from its nest in the trunk of a tree. Checking the surroundings in this manner prior to emerging at dusk for a night of foraging is common behaviour. Found only in Hokkaido, Japan, these animals prefer small holes to their nests, as this provides a measure of protection from predators like owls and martens. This particular nest entrance is also partially covered by a large white fungus, which obscures it from view. This animal is a sub-species of Siberian flying squirrel and is known locally as ezo-momonga.
    japanese-dwarf-flying-squirrel-at-du...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This imdividual was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii), a sub-species of Siberian flying squirrel, emerging from its nest at dusk to set out for a night of foraging. Known locally as ezo-momonga, this sub-species is found only in Hokkaido, Japan. It is primarily nocturnal. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range.
    japanese-dwarf-flying-squirrel-at-du...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii), a sub-species of Siberian flying squirrel, peeking out from its nest before emerging for a night of foraging. Known locally as ezo-momonga, this sub-species is found only in Hokkaido, Japan. It is primarily nocturnal. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range.
    japanese-dwarf-flying-squirrel-at-du...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii) taking a moment to gather itself after emerging from its nest to head out for a night of foraging. Known locally as ezo-momonga, this sub-species of Siberian flying squirrel is found only in Hokkaido, Japan. It is primarily nocturnal. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range.
    japanese-dwarf-flying-squirrel-at-du...tif
  • This is a species of flying squirrel called Ezo Momonga in Japanese (Pteromys volans orii). It is a sub-species of Siberian flying squirrels that is found on the island of Hokkaido in Japan. These squirrels are primarily nocturnal and usually shy, though the squirrels can occasionally be seen during the day. They prefer to take up residence in holes made by woodpeckers. Their diet primarily comprises leaves, seeds, cones, buds, sprouts, nuts and berries. This individual paused in this position for a few moments after emerging one winter evening from its hole in a Sakhalin fir tree (Abies sachalinensis), known as todomatsu in Japanese. This species does not hibernate.
    ezo-momonga-pteromys-volans-orii-fly...tif
  • A pair of Ezo red foxes (Vulpes vulpes schrencki), called Kita Kitsune in Japanese, wandering through a populated area. These foxes are accustomed to the presence of people and can often be seen patrolling areas around houses, farms and other buildings in Hokkaido, Japan.
    vulpes-vulpes-schrencki-ezo-red-foxe...tif
  • This is the Japanese sub-species of Ural owl (Strix uralensis japonica) resting in a tree in Hokkaido. These owls are known as Ezo Fukurou in Japanese.
    strix-uralensis-japonica-ural-owl-ez...tif
  • Lone Japanese crane (Grus japonensis) in a snowstorm. This species is found in Siberia, Northeast China, Mongolia, Korea and northern Japan. The population in northern Japan is mostly non-migratory, remaining resident on the island of Hokkaido throughout the year. This species is listed as Endangered on the IUCN Red List.
    grus-japonensis-red-crowned-japanese...tif
  • This is a Japanese crane (Grus japonensis) in flight in Hokkaido, Japan.
    grus-japonensis-japanese-red-crowned...tif
  • This is a long-tailed tit (Aegithalos caudatus) with winter plumage. Photographed in Hokkaido, Japan.
    long-tailed-tit-aegithalos-caudatus-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This male was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This male was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This male was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • This is a male Japanese dwarf flying squirrel (Pteromys volans orii). This sub-species of Siberian flying squirrel, found only in Hokkaido, Japan and known locally as ezo-momonga, is primarily nocturnal. This male was active during the day because it was competing for the attention of a female in oestrous.
    japanese-dwarf-flying-squirrel-male-...tif
  • Pictured here is a pair of a Japanese dwarf flying squirrel (Pteromys volans orii), one peeking out from the nest, and another perched on a branch above. These squirrels are preparing for a night of foraging, having emerged shortly after sunset. Known locally as ezo-momonga, this sub-species is a  sub-species of Siberian flying squirrel and is found only in Hokkaido, Japan. It is primarily nocturnal. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range.
    japanese-dwarf-flying-squirrels-at-d...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii), a sub-species of Siberian flying squirrel. Known locally as ezo-momonga, this sub-species is found only in Hokkaido, Japan. It is primarily nocturnal. Mature females measure up to 15cm, males up to 18cm (not including tail). These animals weigh up to 120g and are capable of gliding considerable distances. During flight, they use their patagia (membranes of skin between their forelimbs and hind limbs) and tails (10-12cm) to achieve lift, directional control and maneuvering capability. One study in Japan recorded a maximum glide distance exceeding 49m, though most flights fell into the 10m to 20m range. This individual has just emerged shortly after sunset from its nest, a cavity in the tree pictured. After preening for a bit, the flying squirrel ascended into the canopy to forage for the night. While these squirrels are known to nest in a number of tree species, they appear to prefer Sakhalin fir trees (Abies sachalinensis).
    japanese-dwarf-flying-squirrel-at-du...tif
  • This is the Japanese sub-species of Ural owl (Strix uralensis japonica) yawning while sitting in a tree on a snowy day in Hokkaido. These owls are known as Ezo Fukurou in Japanese.
    strix-uralensis-japonica-ural-owl-ya...tif
  • Young Japanese sika deer fawn (Cervus nippon yesoensis) with its tongue out. Photographed in Utoro, Shiretoko National Park, Japan.
    Japanese-sika-deer-fawn-Cervus-nippo...tif
  • This is a Pteromys volans orii flying squirrel consuming male catkins from an Erman's birch tree (Betula ermanii).
    pteromys-volans-orii-flying-squirrel...tif
  • These are Japanese cranes (Grus japonensis) in flight.
    grus-japonensis-japanese-red-crowned...tif
  • This is a pair of Ural owls (Strix uralensis japonica) that have come together at the end of winter in preparation for the spring reproductive season.
    strix-uralensis-japonica-ural-owl-ez...tif
  • This is a female Strix uralensis japonica owl perched at the entrance to her nest during snowfall. A smaller male was inside the nest.
    strix-uralensis-japonica-ural-owl-ez...tif
  • This is an Ezo chipmunk (Tamias sibiricus lineatus), recently emerged from winter hibernation.
    tamias-sibiricus-lineatus-ezo-chipmu...tif
  • This is a Pteromys volans orii flying squirrel bathed in the cool light of an early winter morning. These animals are normally nocturnal.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Japanese dwarf flying squirrel (Pteromys volans orii) flying through the forest on a cold winter day. Usually nocturnal, these animals sometimes become active in the day during reproductive season.
    japanese-flying-squirrel-pteromys-vo...tif
  • This is a male Pteromys volans orii flying squirrel sitting in a favorite spot on a sunny day, looking down upon his surroundings.
    pteromys-volans-orii-flying-squirrel...tif
  • This Pteromys volans orii flying squirrel has carried a few sticks of Japanese elm (Ulmus davidiana var. japonica) back to its nest at the end of foraging to enjoy a takeaway meal before going inside.
    pteromys-volans-orii-flying-squirrel...tif
  • This is a Pteromys volans orii flying squirrel flying through trees.
    pteromys-volans-orii-flying-squirrel...tif
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Tony Wu Underwater Photography

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