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  • I photographed this Pachyseris speciosa coral formation in Disney Lake, a saltwater enclosure in the Rock Islands of Palau. The lake is only accessible via a submerged tunnel. The lake and surrounding area is sheltered from wind and waves, and almost no one enters the enclosed area, so some species of coral that are normally fragile are able to grow to large sizes. This coral formation was so thin at its edge that light can shine through. Lit from behind, the lines comprising the coral growth reminded me of flowing volcanic lava. See separate close-up photograph of coral edge detail.
    pachyseris-speciosa-coral-disney-lak...tif
  • I photographed this Pachyseris speciosa coral formation in Disney Lake, a saltwater enclosure in the Rock Islands of Palau. The lake is only accessible via a submerged tunnel. The lake and surrounding area is sheltered from wind and waves, and almost no one enters the enclosed area, so some species of coral that are normally fragile are able to grow to large sizes. This coral formation was so thin at its edge that light can shine through. Lit from behind, the lines comprising the coral growth reminded me of flowing volcanic lava.
    pachyseris-speciosa-coral-disney-lak...tif
  • Healthy cabbage coral (Turbinaria reniformis) formation surrounded by basslets, damsels and other tropical reef fish. Photographed at about 15 meters depth on a dive site near Normanby Island in Milne Bay, Papua New Guinea that we named Julian's Cabbage Patch because there was so much cabbage coral.
    cabbage-coral-turbinaria-reniformis-...tif
  • This is a close-up view of leaf coral (Pavona decussata) spawning, with streams of sperm being sent into the water column in synchrony with gametes from other nearby colonies of the hard coral.
    pavona-deccusata-leaf-coral-spawning...tif
  • This is a colony of Lobophyllia corymbosa hard coral engaged in spawning. Hard corals like this release both eggs and sperm at the same time, as seen here.
    lobophyllia-corymbosa-hard-coral-spa...tif
  • Depicted here is spawning of a leaf coral (Pavona decussata) formation. Spawning was synchronized over a large area with many coral colonies.
    pavona-decussata-leaf-coral-spawning...tif
  • Cute yellow coral goby (Gobiodon okinawae) living among branches of Acropora coral at Fantasy Island, part of the Patch Reefs of Palau. This area was devastated by the el Nino of 1998 and a subsequent crown of thorns infestation. The area has since recovered and is a thriving, healthy reef system.
    yellow-coral-goby-gobiodon-okinawae-...tif
  • Fusilier damselfish (Lepidozygus tapeinosoma) swarming over large staghorn coral (Acropora sp.) formation at Carl's Ultimate dive site in the Eastern Fields of Papua New Guinea.
    fusilier-damselfish-Lepidozygus-tape...tif
  • Red-tinted black coral tree (Antipathes sp.) surrounded by a school of orange fairy basslets (Pseudanthias squamipinnis) at the Black and Silver dive site in the Nuakata Island area of Milne Bay, Papua New Guinea.
    Antipathes-sp-black-coral-Pseudanthi...tif
  • Multicolored soft corals inflated to feed in the current passing over Sullivan's Patches, a beautiful coral reef located in Milne Bay, Papua New Guinea
    multicolored-soft-coral-sullivans-pa...tif
  • Cute coral crab (Trapezia sp.) in pretty mint-colored hard coral (Acropora sp.)
    cute-Trapezia_crab-in-coral-ambon-20..tiff
  • This is a colony of Lobophyllia hemprichii hard coral engaged in boadcast spawning. Some eggs are visible on the left side of the image, floating through streams of sperm. Nearby colonies of the same species released gametes in synchrony.
    lobophyllia-hemprichii-coral-broadca...tif
  • This is a colony of leaf coral (Pavona decussata) spawning, with streams of sperm being sent into the water column in synchrony with gametes from other nearby colonies of the hard coral.
    pavona-deccusata-leaf-coral-spawning...tif
  • This is a coral grouper (Cephalopholis miniata) pictured with a typical reef scene in the Japanese island of Kumejima in Okinawa.
    cephalopholis-miniata-coral-grouper-...tif
  • A lush, colourful coral formation with soft and hard coral, sea fans and a school of bright-orange fairy basslets at Carl's Ultimate dive site in the Eastern Fields of Papua New Guinea
    coral-formation-jewel-fairy-basslet-...tif
  • This is a polyp of Alveopora japonica, a small zooxanthellate scleractinian coral that produces large polyps. The orange spheres are planulae, or larval corals. Individual planula migrate from the coral stalk to the mouth, where the planula emerges and swims into the water. The species is found in Japan, Korea and Taiwan. Recent research suggests that genetic differences among the three populations are large enough to warrant consideration as separate species.
    alveopora-japonica-coral-planula-jap...tif
  • Staghorn coral formation at Point P dive site in the Eastern Fields of Papua New Guinea, with a large school of bigeye trevallies (Caranx sexfasciatus) swimming overhead
    caranx-sexfasciatus-bigeye-trevally-...tif
  • Cute yellow coral goby living among branches of Acropora coral at Fantasy Island, part of the Patch Reefs of Palau. This area was devastated by the el Nino of 1998 and a subsequent crown of thorns infestation. The area has since recovered and is a thriving, healthy reef system.
    yellow-coral-goby-gobiodon-okinawae-...tif
  • Pontonides sp. commensal shrimp on a curled-up whip coral (Cirrhipathes sp.) formation
    Pontonides-commensal-shrimp-on-Cirrh...tif
  • Plush strawberry sherbet red Dendronephthya sp. soft coral with lots of fish swimming in the background. Photographed at Surprise Reef near Nuakata Island in Milne Bay, Papua New Guinea.
    Dendronephthya-sp-red-soft-coral-mil...tif
  • Large, unspoiled staghorn coral (Acropora sp.) viewed from the bottom-up, photographed at Craig's Ridge dive site in Moresby Strait, Milne Bay, Papua New Guinea
    Acropora-sp-coral-milne-bay-papua-ne...tif
  • Thousands of Chromis damselfish and Pseudanthias basslets swarming over a healthy Acropora hard coral formation in shallow water at Ashmore Reef in Australian waters.
    Acropora-hard-coral-with-thousands-o...tif
  • Diver with an illuminated plush pink soft coral growing in the muck of Lembeh Strait, Indonesia
    pink-soft-coral-diver-lembeh-strait-...tif
  • Shown here are oriental butterflyfishes (Chaetodon auripes) consuming gametes from spawning Lobophyllia hemprichii hard coral. Several species of reef fish engage in this type of opportunistic feeding. Photographed in Kagoshima Prefecture, Japan.
    chaetodon-auripes-butterflyfish-eati...tif
  • This is a polyp of Alveopora japonica, a small zooxanthellate scleractinian coral that produces large polyps. The orange spheres are planulae, or larval corals. Individual planula migrate from the coral stalk to the mouth, where the planula emerges and swims into the water. The species is found in Japan, Korea and Taiwan. Recent research suggests that genetic differences among the three populations are large enough to warrant consideration as separate species.
    alveopora-japonica-coral-planula-jap...tif
  • This is a colony of Pavona decussata leaf coral, photographed one day prior to spawning.
    pavona-decussata-leaf-coral-japan-20...tif
  • Whip coral shrimp (Pontonides ankeri) on a sea whip (Cirripathes sp.) in Lembeh Strait, Indonesia. This species is often confused with Pontonides unciger, which lives on bushy black corals (Anthipathes sp.)
    pontonides-ankeri-shrimp-on-cirripat...tif
  • Shallow, protected nooks and crannies among the Rock Islands of Palau are characterized by murky water, but can provide a safe environment for corals to flourish and develop into intricate, unbroken formations like the one pictured here. This coral tree was one of many in very shallow water in front of Ngchuus Beach, Ngeruktabel Island.
    backlit-coral-formation-ngeruktabel-...tif
  • Very small blackfin coral goby (Paragobiodon lacunicolus) hiding among the branches of a small coral formation in the muck at Lembeh Strait, North Sulawesi, Indonesia
    blackfin-coral-goby-Paragobiodon-lac...tif
  • Coral and sponge formation on the outer reef in Ambon, Indonesia
    soft-coral-sea-fan-sponge-ambon-indo...tif
  • Plush, healthy soft coral formation on the reef in Ambon, Indonesia
    healthy-soft-coral-formation-ambon-i...tif
  • Colorful Dendronephthya sp. soft coral with fish swimming in the background. Photographed at Surprise Reef near Nuakata Island in Milne Bay, Papua New Guinea.
    colorful-Dendronephthya-soft-coral-n...tif
  • Red coral formation at 40 metres depth, with silhouette of lone grey reef shark patrolling in the background
    red-coral-grey-reef-shark-ashmore-re...tif
  • This is a polyp of Alveopora japonica, a small zooxanthellate scleractinian coral that produces large polyps. The orange spheres are planulae, or larval corals. Individual planula migrate from the coral stalk to the mouth, where the planula emerges and swims into the water. The species is found in Japan, Korea and Taiwan. Recent research suggests that genetic differences among the three populations are large enough to warrant consideration as separate species.
    alveopora-japonica-coral-planula-jap...tif
  • Diver Terry Ward shining a light on multicolored Lobophyllia brain coral formation at Darwin's Wall that Ron Leidich has dubbed the Four Corners. This unique location is flushed by strong currents exchanged between a large inner lake (Long Lake) and the open ocean, generating ample nutrient flow to support these slow-growing corals, which flourish in large numbers and grow to exceptional size in this area. There is minimal sunlight that penetrates the canopy of overhanging foliage, meaning than the corals rely more on the daily tidal exchange for nutrients than on their zooxanthallae. This wall and the corals that live on them are a perfect illustration of the intricate relationship between the ecosystems of the inner lakes of Palau's Rock Islands and the open ocean.
    diver-with-lobophyllia-coral-darwins...tif
  • Large, beautiful black coral (Anthipathes sp.) formation at Hukurila Cave in Ambon, Indonesia
    Anthipathes-black-coral-ambon-indone...tif
  • Millions of fusilier damselfish (Lepidozygus tapeinosoma) swarming over a pristine coral reef called Carl's Ultimate, located in the Eastern Fields of Papua New Guinea
    pristine-coral-reef-millions-of-fish...tif
  • The marine lakes of Palau comprise a plethora of hidden environments and fragile ecosystems, often characterized by delicate formations such as this Merulina sp. hard coral, which was thin enough to be translucent. Photographed inside Disney Lake, located in Nikko Bay, Palau.
    translucent-merulina-sp-hard-coral-d...tif
  • The marine lakes of Palau comprise a plethora of hidden environments and fragile ecosystems, often characterized by delicate formations such as this Merulina sp. hard coral, which was thin enough to be translucent. Photographed inside Disney Lake, located in Nikko Bay, Palau.
    translucent-merulina-sp-hard-coral-d...tif
  • Soft coral on the outer reef of Ambon, Indonesia
    soft-coral-ambon-indonesia-200810-00...tif
  • Wire coral goby (Bryaninops yongei) photographed with a bit of motion-blur to show this fish opening its mouth
    whip-coral-goby-Bryaninops-yongei-mo...tif
  • The steep, conical shape of this Porites sp. hard coral formation is reminiscent of the rock islands that comprise the land mass of Palau.
    porites-sp-hard-coral-formation-pala...tif
  • Large Platygyra hard coral in shallow blue water in tropical water of Indonesia
    Spherical-Playtgyra-coral-Ambon-2010..tiff
  • This is a Dendronephthya soft coral colony growing atop a rock, where currents provide ample nutrient flow. The light in the background is from a scuba diver.
    dendronephthya-soft-coral-japan-2022...tif
  • Phyllodesmium jakobsenae nudibranch feeding on Xenia soft coral in Lembeh Strait, North Sulawesi, Indonesia
    Phyllodesmium-jakobsenae-nudibranch-...tif
  • Coral hawkfish (Cirrhitichthys oxycephalus) perched on a sponge, using its pectoral fins for balance. Lembeh Strait, North Sulawesi, Indonesia.
    coral-hawkfish-Cirrhitichthys-oxycep...tif
  • Large Platygyra hard coral in shallow blue water in tropical water of Indonesia
    Platygyra-coral-against-blue-water-a..tiff
  • Coral cardinalfish (Siphamia corallicola) with a mouthful of orange eggs
    Siphamia-corallicola-coral-cardinalf...tif
  • This is close-up view of polyps of Alveopora japonica, a small zooxanthellate scleractinian coral that produces large polyps. This species is found in Japan, Korea and Taiwan. Recent research suggests that genetic differences among the three populations are large enough to warrant consideration as separate species.
    alveopora-japonica-coral-polyps-japa...tif
  • Bright red sabre squirrelfish (Sargocentron spiniferum) peeking out from underneath a large, healthy coral formation on Carl's Ultimate dive site in the Eastern Fields of Papua New Guinea
    sabre-squirrelfish-sargocentron-spin...tif
  • Red and orange soft coral (Dendronephthya sp.) in the Eastern Fields of Papua New Guinea, framed by a curtain of thousands of bigeye trevallies (Caranx sexfasciatus)
    soft-coral-bigeye-trevally-Caranx-se...tif
  • The corals living in the shallow waters around Fantasy Island in the Patch Reefs of Palau were devastated by a massive crown of thorns attack following extensive bleaching damage caused by El Niño in 1998. This photograph was taken in 2011, just 13 years after the corals here were wiped out.
    healthy-coral-recovered-bleaching-cr...tif
  • Healthy reef systems grow continuously, as illustrated here, with new coral growth atop old coral formations. Robust old-growth coral formations like this are common in areas without much human activity, such as the Eastern Fields of Papua New Guinea.
    robust-coral-growth-eastern-fields-p...tif
  • Soft coral with lots of magenta slender Anthias and a couple of golden damselfish, on a steep wall in the Eastern Fields of Papua New Guinea
    soft-coral-wall-dive-eastern-fields-...tif
  • When the tides change and the currents run at Carl's Ultimate dive site in the Eastern Fields of Papua New Guinea, fish of all description swarm the reef by the millions. I've seen more fusilier damselfish (Lepidozygus tapeinosoma) like the ones pictured here at this dive site than I have at any other location I've ever dived. It is truly one of the world's greatest spectacles.
    Luzonichthys-waitei-basslets-swarmin...tif
  • Two Paguritta coral hermit crabs looking out at the world from their burrows in a coral formation.
    Paguritta-coral-hermit-crabs-eastern...tif
  • Spectacular pink and orange soft corals in full bloom, surrounded by multicoloured fairy basslets
    pink-and-orange-soft-coral-picasso-p...tif
  • A lone sponge perched on a ledge in the Eastern Fields, a submerged coral reef atoll in the Coral Sea of Papua New Guinea. Photographed to evoke the image of a solitary sentinel watching over the deep sea.
    illuminated-marine-sponge-coral-sea-...tif
  • Plush Dendronephthya orange soft corals at Sahaung 1 dive site at Bangka Island, North Sulawesi, Indonesia
    orange-Dendronephthya-soft-coral-ban...tif
  • A swarm comprising thousands of orange basslets (Pseudanthias squamipinnis) rising up from an enormous hard coral formation at the top of Lynda's Reef near Nuakata Island in Milne Bay, Papua New Guinea
    Pseudanthias-squamipinnis-swarm-lynd...tif
  • Solitary longfin bannerfish swimming along a coral reef. This fish appeared to be patrolling a limited area, with a tendency to return to this one spot. After a while, the fish grew accustomed to my presence and approached closely.
    longfin-bannerfish-pennant-coralfish...tif
  • A gigantic, unbroken sea fan with swarming Anthias at Ashmore Reef in Australian waters in the Coral Sea. The first time we dived this site, I fell in love with the area because there were so many large, unspoiled sea fans like this one, and the reef was teeming with millions of small fish. Captain Craig DeWit of MV Golden Dawn was nice enough to name the dive site Wu Wu Wall for me.
    sea-fan-anthias-basslets-wu-wu-wall-...tif
  • A beautiful Lori's Anthias (Pseudanthias lori). This fish is male, with characteristic red, orange, yellow, blue and purple coloration on its body. Photographed at Carl's Ultimate dive site in the Eastern Fields of Papua New Guinea, a submerged coral reef atoll in the Coral Sea
    Pseudanthias-lori-basslet-anthias-ea...tif
  • This is a red-spotted blenny (Blenniella chrysospilos) poking its head out of its home in the coral, with two coral hermit crabs (Paguritta sp.) visible in the background. The blenny's home is the abandoned burrow of a vermetid gastropod, Dendropoma maximum.
    blenniella-chrysospilos-red-spotted-...tif
  • A beautiful Lori's Anthias (Pseudanthias lori). This fish is male, with characteristic red, orange, yellow, blue and purple coloration on its body. Photographed at Carl's Ultimate dive site in the Eastern Fields of Papua New Guinea, a submerged coral reef atoll in the Coral Sea
    colorful-Pseudanthias-lori-basslet-m...tif
  • A beautiful male Lori's Anthias (Pseudanthias lori), with characteristic red, orange, yellow, purple coloration on its body. Photographed at Carl's Ultimate dive site in the Eastern Fields of Papua New Guinea, a submerged coral reef atoll in the Coral Sea
    Pseudanthias-lori-anthias-basslet-ea...tif
  • Orange-yellow crinoid feather star perched on a coral branch, filtering nutrients from the passing current. The position and color of this crinoid reminded me of a sunflower.
    crinoid-feather-star-with-backlighti...tif
  • Pretty pink Commerson's frogfish sitting on a sponge concealed by sea fans, at a dive site called Shaw Thing in the Eastern Fields of Papua New Guinea. This dive was the first time that Captain Craig of Golden Dawn had come across one of these frogfish in the Eastern Fields even though he had been diving the area for about 20 years at the time of this encounter.
    pretty-pink-frogfish-Antennarius-com...tif
  • There are still virgin reefs like this that few people have seen, where unspoiled corals carpet the reef system and countless fish gather to feed in the currents. This photograph is from a dive site called Tokyo Express in the Eastern Fields of Papua New Guinea.
    unspoiled-coral-reef-millions-of-fis...tif
  • Hardyhead silversides (Atherinomorus lacunosus) swarming around the coral-encrusted remains of a portion of the jetty at Samarai Island in Milne Bay, Papua New Guinea
    Hardyhead-silversides-Atherinomorus-...tif
  • Thousands of hardyhead silversides (Atherinomorus lacunosus) circling around the coral-encrusted remains of a jetty post at Samarai Island in Milne Bay, Papua New Guinea.
    Hardyhead-silversides-Atherinomorus-...tif
  • Lionfish patrolling the edge of Suzie's Bommie, a popular dive site accessible from Lololata Island Resort near Port Moresby. This small but healthy reef is always teeming with life, including large schools of sweetlips, thousands of basslets and many other marine organisms. This image is licensed on an exclusive basis to BBC through 29 October 2021.
    lionfish-Pterois-volitans-loloata-su...tif
  • Beautiful, undamaged sea fans like this are characterstic of healthy tropical reefs. Photograph taken in the Eastern Fields of Papua New Guinea
    sea-fan-and-evening-sun-eastern-fiel...tif
  • When small fish like basslets, damselfish and other reef residents appear in large numbers with the change of tide at Carl's Ultimate dive site in the Eastern Fields of Papua New Guinea, predators such as bluefin trevally gather in packs to hunt them. This trevally used me for camouflage to approach the swarms of small fish, then charged straight in front of me in an attempt to grab a meal.
    bluefin-trevally-Caranx-melampygus-h...tif
  • Unspoiled orange-yellow sea fan with golden damselfish and orange basslets in the Eastern Fields of Papua New Guinea
    orange-yellow-sea-fan-anthias-damsel...tif
  • Strong currents around Bangka Island bring nutrients to support sea fans like this. Note the small group of juvenile convict fish at the lower left part of the sea fan. Juvenile convict fish mimic venemous striped catfish (Plotosus lineatus). These fish were mistakenly called convict blennies and convict gobies, but are neither blennies nor gobies.
    red-sea-fan-blue-water-juvenile-conv...tif
  • Healthy gorgonian sea fans at Deacon's Reef in Milne Bay, Papua New Guinea. Bob Halstead has picked up broken sea fans over many years and planted these sea fans on this wall, producing a beautiful dive site, with overhanging trees visible above.
    red-gorgonian-sea-fan-deacons-reef-m...tif
  • Wall of gorgonian sea fans at Deacon's Reef in Milne Bay, Papua New Guinea. Bob Halstead has picked up broken sea fans over many years and planted these sea fans on this wall, producing a beautiful dive site, with overhanging trees visible above.
    red-gorgonian-sea-fan-deacons-reef-m...tif
  • Orange basslets with sea fans on top of the arch at The Doghouse dive site in Diving Dog Passage on the Barrier Reef of Papua New Guinea
    Pseudanthias-squamipinnis-orange-bas...tif
  • Orange basslets with sea fans on top of the arch at The Doghouse dive site in Diving Dog Passage on the Barrier Reef of Papua New Guinea
    Sea-fan-with-Pseudanthias-squamipinn...tif
  • Large sea fan with magenta slender basslets, Eastern Fields, Papua New Guinea. This photograph is a film scan.
    sea-fan-eastern-fields-papua-new-gui...tif
  • Large group of sea fans with lots of jewel fairy basslets, at Carl's Ultimate dive site in the Eastern Fields of Papua New Guinea
    sea-fans-with-jewel-fairy-basslets-P...tif
  • Pseudanthias squamipinnis orange basslets with sea fans on top of the arch at The Doghouse dive site in Diving Dog Passage on the Barrier Reef of Papua New Guinea
    Orange-basslets-Pseudanthias-squamip...tif
  • Diver with large, healthy Tridacna gigas giant clam at a remote location in Papua New Guinea
    tridacna-gigas-giant-clam-and-diver-...tif
  • Lubricogobius dinah goby sitting on top of its beer bottle home, located at 30 meters depth at Observation Point at Normanby Island in Milne Bay province, Papua New Guinea.
    Lubricogobius-dinah-goby-observation...tif
  • Colorful male jewel fairy basslet (Pseudanthias squamipinnis) photographed in the Eastern Fields of Papua New Guinea
    jewel-fairy-basslet-male-eastern-fie...tif
  • Enormous school of hardyhead silversides (Atherinomorus lacunosus) swarming under the jetty at Samarai Island in Milne Bay, Papua New Guinea.
    school-of-hardyhead-silversides-Athe...tif
  • Pair of Dinah's gobies (Lubricogobius dinah) with their beer-bottle home, at Observation Point in Milne Bay Province, Papua New Guinea
    Dinahs-goby-pair-Lubricogobius-dinah...tif
  • Darkfin chromis (Chromis atripes), photographed in the Eastern Fields of Papua New Guinea
    darkfin-chromis-atripes-papua-new-gu...tif
  • Super macro close-up 3x life-size magnification photograph of a rayed shrimp goby (Tomiyamichthys sp.), taken at Waga Waga in Milne Bay, Papua New Guinea
    rayed-shrimp-goby-Tomiyamichthys-sp-...tif
  • Countless robust silversides (Atherinomorus lacunosus) swarming under the jetty at Samarai Island in Milne Bay, Papua New Guinea.
    school-of-robust-silversides-Atherin...tif
  • School of Yellowstripe Scad (Selaroides leptolepis) swimming under the jetty at Samarai Island in Milne Bay, Papua New Guinea
    Selaroides-leptolepis-yellowstripe-s...tif
  • School of Yellowstripe Scad (Selaroides leptolepis) swimming under the jetty at Samarai Island in Milne Bay, Papua New Guinea, one fish with its mouth wide open.
    Selaroides-leptolepis-yellowstripe-s...tif
  • Aegires gardineri nudibranch photographed at Sawa Sawa Sawaga in Milne Bay, Papua New Guinea
    Aegires-gardineri-nudibranch-milne-b...tif
  • Dinah's goby (Lubricogobius dinah) sitting on top of a beer bottle. Observation Point at Normany Island in Milne Bay province, Papua New Guinea.
    Lubricogobius-dinah-goby-observation...tif
  • Bright tangerine-orange lacy scorpionfish in Papua New Guinea
    tangerine-colored-rhinopias-aphanes-...tif
  • Jewel fairy basslet in Milne Bay, Papua New Guinea. This photograph is a film scan.
    threadfin-anthias-pseudanthias-squam...tif
  • Colourful fairly basslets and sea fans snatching bits of food from the passing current
    fairy-basslets-sea-fan-bangka-north-...tif
  • Discarded plastic garbage on a beach on tourist destination Lizard Island, Australia
    garbage-australia-201206-0184.tif
  • Head-on view of a male jewel fairy basslet (Pseudanthias squamipinnis), photographed in the Eastern Fields of Papua New Guinea
    jewel-fairy-basslet-pseudanthias-squ...tif
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