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  • Atlantic walrus (Odobenus rosmarus rosmarus) sitting on ice, Svalbard.
    atlantic-walrus-on-ice-svalbard-2015...tif
  • A pair of Atlantic walruses (Odobenus rosmarus rosmarus) hauled out on ice during the Arctic summer
    odobenus-rosmarus-atlantic-walruses-...tif
  • Four Atlantic walruses (Odobenus rosmarus rosmarus) hanging out in shallow water in Svalbard.
    four-atlantic-walruses-odobenus-rosm...tif
  • Atlantic puffin (Fratercula arctica) perched at the edge of a cliff early in the morning.
    atlantic-puffin-on-cliff-iceland-fra...tif
  • Five Atlantic puffins (Fratercula arctica) perched on a cliff edge in Iceland, bathed by warm light of Arctic summer sunset.
    five-atlantic-puffins-on-cliff-sunse...tif
  • A pair of Atlantic puffins (Fratercula arctica) in warm sunlight prior to sunset in the Arctic summer. The puffin in the rear is stretching and flapping its wings.
    atlantic-puffins-sunset-iceland-frat...tif
  • Atlantic puffin (Fratercula arctica) scratching its head
    atlantic-puffin-scratching-head-icel...tif
  • A pair of Atlantic puffins (Fratercula arctica) socializing late at night, just before the Arctic summer sunset in Iceland.
    atlantic-puffins-socializing-sunset-...tif
  • Atlantic puffin (Fratercula arctica) at sunrise, Iceland
    atlantic-puffin-sunrise-iceland-frat...tif
  • Atlantic puffin (Fratercula arctica) basking in the warm light of winter sunrise, Iceland
    atlantic-puffin-fratercula-arctica-i...tif
  • Atlantic puffin (Fratercula arctica) approaching cliff for a landing in the middle of the night.
    atlantic-puffin-landing-on-cliff-fra...tif
  • Atlantic puffin (Fratercula arctica) swimming on the ocean surface after divind down to catch fish
    atlantic-puffin-fratercula-arctica-s...tif
  • Atlantic puffin (Fratercula arctica) shaking off water after returning from sea
    atlantic-puffin-shaking-off-water-fr...tif
  • Atlantic puffin (Fratercula arctica) collecting nesting materials at night in the Artic summer of Iceland.
    atlantic-puffin-gathering-nesting-ma...tif
  • Atlantic puffin (Fratercula arctica) with its mouth open.
    atlantic-puffin-open-mouth-fratercul...tif
  • Profile of an Atlantic puffin (Fratercula arctica) in Iceland
    atlantic-puffin-face-fratercula-arct...tif
  • Atlantic Puffin (Fratercula arctica) with a mouthful of nesting material pulled from the ground near its burrow. Photographed in Iceland
    atlantic-puffin-mouthful-nesting-mat...tif
  • Group of Atlantic walruses (Odobenus rosmarus rosmarus) resting on ice in Svalbard
    walrus-group-on-ice-svalbard-odobenu...tif
  • Atlantic walrus (Odobenus rosmarus rosmarus) sleeping on ice during a sunny summer evening in Svalbard.
    sleeping-walrus-odobenus-rosmarus-sv...tif
  • Four Atlantic walruses (Odobenus rosmarus rosmarus) relaxing on a beach in Svalbard
    four-walruses-relaxing-on-beach-sval...tif
  • Young walrus (Odobenus rosmarus) with small, undamaged tusks
    atlantic-walrus-odobenus-rosmarus-ro...tif
  • Puffins (Fratercula arctica) basking in warm, red sunlight of dawn during the Arctic summer season in Iceland.
    atlantic-puffins-basking-red-dawn-su...tif
  • This is a north Atlantic humpback whale (Megaptera novaeangliae novaeangliae) pursuing a large school of herring in the cold, dark water of Arctic winter. The humpback whale swallowed several mouthfuls of fish during this foraging period.
    humpback-whale-pursuing-herring-norw...tif
  • Young Atlantic walrus (Odobenus rosmarus rosmarus) swimming underwater, photographed in Svalbard
    young-walrus-underwater-svalbard-odo...tif
  • A group of Atlantic walruses (Odobenus rosmarus rosmarus) swimming rapidly while foraging for food in shallow water late at night, with one raising its head high to take a look around
    walrus-spyhop-while-foraging-svalbar...tif
  • A walrus (Odobenus rosmarus) demonstrating use of its tusks. There were no serious injuries resulting from this inter-walrus discussion. Both individuals settled down and went back to sleep.
    walrus-stabbing-another-walrus-with-...tif
  • Walrus (Odobenus rosmarus) with a runny nose, sound asleep on ice in Svalbard
    sleeping-walrus-with-runny-nose-sval...tif
  • Walruses (Odobenus rosmarus) resting on ice, with two large individuals facing off just before the one on the right stabbed the other with his tusks.
    walruses-on-ice-facing-off-svalbard-...tif
  • Walrus (Odobenus rosmarus) with a runny nose sleeping on ice, flipper covering his head
    sleeping-walrus-runny-nose-svalbard-...tif
  • Large aggregation of walruses (Odobenus rosmarus) socializing late at night and foraging for food. This chaotic and noisy group activity continued for many hours, with several dozen individuals involved. The large group sometimes broke up into two or three smaller groups.
    walrus-social-group-foraging-svalbar...tif
  • A large colony of walruses (Odobenus rosmarus rosmarus) hauled up on a sandy beach area to rest. Individuals occasionally ventured into the water for a while before returning to the group to sleep.
    sleeping-walrus-colony-svalbard-norw...tif
  • This is the internal shell of a deep-water cephalopod called a Ram’s horn squid, also known as a little post horn squid (Spirula spirula). This cephalopod is the only known member of its genus. The internal shell is both robust and buoyant, so often found on beaches. This one washed up during a mass stranding of thousands of Physalia utriculus amd other animals of the open-ocean, pelagic community. Attached to the shell were pelagic goose barnacles (Lepas sp.).
    rams-horn-shell-spirula-goose-barnac...tif
  • Top-down view of an Indo-Pacific Portuguese Man-of-War (Physalia utriculus). This is one of many thousands that were part of a mass stranding in South Africa. Such strandings happen from time to time, bringing these colonial siphonophores to shore, along with other associated animals such as Velella velella, Janthina janthina, and Planes major from the rarely seen open-ocean, blue-water community of life. The beautiful blue and green skirt surrounding the air pocket (pneumatophore) are loaded with nematocysts, which deliver potent venom
    blue-bottle-indo-pacific-portuguese-...tif
  • Many orcas passing at depth. The shallowest orca was at about 10m. There were many more scattered in all directions.
    orcas-swimmng-at-depth-norway-201601...tif
  • Female polar bear (Ursus maritimus) with a single young cub, only a few months old. Polar bears usually have two cubs. It is possible that this female lost a cub. Photographed in northern Svalbard.
    polar-bear-female-and-cub-svalbard-u...tif
  • Pandemonium and commotion among a group of socializing sperm whales (Physeter macrocephalus)
    sperm-whale-social-group-physeter-ma...tif
  • This is a horned isopod (Deto echinata), a species of air-breathing isopod that inhabits seashores of southern Africa. This isopod is a woodlouse, which is an isopod that is adapted to terrestrial rather than aquatic life. This species lives onshore, feeding in the intertidal zone on carrion and plant material washed up by the ocean, as well as small live prey. The horns on the backs of this species are longer in males than in females. This is a male, about two centimeters in length.
    horned-isopod-deto-echinata-201711-3...tif
  • This is a violet snail (Janthina janthina), also known as a bubble raft snail. These pelagic snails are found worldwide in temperate and tropical waters. They are considered pleuston, macroscopic organisms that live at the interface of air and water. To float at the ocean surface, these snails use thin secretions of chitin to envelope bubbles of air, thus creating a buoyancy support. They bob upside-down below the ocean surface and wait for prey such as Portuguese man-of-wars (Physalia sp.) and by-the-wind sailors (Velella velella). When they sense such prey, bubble raft snails extend their long cylindrical snouts and use a rasping mechanism to grab prey, as pictured here. These snails are protandrous sequential hermaphrodites, meaning they all commence life as males, then become female. This snail, with a shell of about three centimeters, was found washed ashore with a mass stranding of thousands of colonial siphonophores.
    janthina-janthina-violet-bubble-raft...tif
  • This is a parasite that was associated with a Mola mola ocean sunfish. It is a copepod of the family Caligidae (possibly Caligus sp. or Lepeoptheirus sp.), the carapace slightly over 1cm in length. The long appendage visible at the rear end is one of two original appendages. The other one fell off. The one that is visible is actually split into two. Each slit carried a developing juvenile copepod. There is one remaining one visible at the distal end.
    parasitic-copepod-associated-with-mo...tif
  • Close-up of a sunfish (Mola mola). This individual was floating on the ocean surface, relaxed and calm. The fish approached the boat numerous times, allowing me on one occasion to remove one of the copepod parasites visible in this photograph. When I entered the water and floated nearby, the fish bumped directly into me, resulting in this close encounter.
    mola-mola-sunfish-close-encounter-so...tif
  • Mature male killer whale (Orcinus orca), with his tall dorsal fin in full display, swimming in the cold waters of Norway
    mature-male-orca-norway-201601-0566.tif
  • Sperm whale (Physeter macrocephalus) social unit engaged in social activity, with one whale separated from the group while passing gas. Flatulence is common when sperm whales socialize near the ocean surface.
    sperm-whale-flatulence-socializing-d...tif
  • Large adult male killer whale (Orcinus orca) stalking a large school of herring in shallow water.
    orca-hunting-herring-norway-201601-0...tif
  • Mature male orca (Orcinus orca) in the foreground, with other mature males visible in the background. Pairs and small groups of mature males were common in this area of Norway, swimming apart from their larger social units. Mature males are easily recognizable by their prominent dorsal fins.
    orca-adult-male-norway-orcinus-orca-...tif
  • Portrait of sperm whale's enlarged forehead, which contains the spermaceti that was the reason for large-scale hunting of these animals. This individual's name is Scar, an unusually friendly male sperm whale that was about ten years old at this encounter. This image received a Highly Commended recognition in the Gerald Durrell Award for Endangered Wildlife category at the 2010 Veolia Environnement Wildlife Photographer of the Year competition (organised by BBC Wildlife Magazine and the Natural History Museum London)
    scar-male-sperm-whale-dominica-20100...tif
  • Group photo: Silhouettes against ice, created by late evening Arctic sun in Svalbard. Jon Cornforth, Nana Trongratanawong, Colin Lee, Tony Wu, Jenny Huang (left to right)
    group-photo-silhouette-arctic-summer...tif
  • Arctic tern (Sterna paradisaea) hovering above the ocean before it dived in to catch fish.
    arctic-tern-hovering-above-ocean-hun...tif
  • Polar bear (Ursus maritimus) feeding on the remains of a beluga whale (Delphinapterus leucas), with a seagull waiting for its turn. Multiple bears visited this carcass over many days.This bear was wearing a large radio tracking collar, not visible from this angle.
    polar-bear-foraging-beluga-whale-car...tif
  • A very young polar bear cub (Ursus maritimus), probably only a few months old, standing up to take a look around. This cub was alone with its mother. Given that polar bears generally have two cubs, it is possible that something happened to this cub’s sibling. Photographed in Svalbard.
    polar-bear-cub-ursus-maritimus-svalb...tif
  • Close-up view of the ventral surface of an adult female sperm whale (Physeter macrocephalus), with three others in the background.
    sperm-whales-physeter-macrocephalus-...tif
  • Four sperm whales (Physeter macrocephalus) hanging head down in the water, engaged in social interaction. Sperm whales are tactile, often making body contact with other whales, as shown here. They also produce a lot of sound during social interaction like this. Photo taken in Roseau, Dominica.
    four-sperm-whales-socializing-hangin...tif
  • Portrait of a mature female sperm whale (Physeter macrocephalus)
    sperm-whale-physeter-macrocephalus-d...tif
  • Rising tide in front of the Dalebrook tidal pool in Kalk Bay, South Africa
    dalebrooke-tidal-pool-kalk-bay-south...tif
  • This is a Columbus crab (Planes major), a small oceanic crab that lives on floating objects such as seaweed, driftwood and other debris, or in association with animals such as by-the-wind sailors (Velella velella) or turtles. They can appear in different colors. This one was found together with a mass stranding of thousands of Portuguese man-of-war colonial siphonophores (Physalia utriculus) and by-the-wind sailors. The crab was between one and two centimeters. This species was previously known by the name Planes cyaneus.
    columbus-crab-planes-major-south-afr...tif
  • Close encounter with a friendly sunfish (Mola mola) that was floating on the ocean surface, relaxed and calm. The fish approached the boat numerous times, allowing me on one occasion to remove one of the many copepod parasites visible in this photograph. Mola mola are known to host multiple species of parasites, including copepods and nematodes. The fish is floating on one side, with its mouth open, facing the camera.
    sunfish-mola-mouth-open-with-parasit...tif
  • Profile view of a mature male killer whale (Orcinus orca) in the cold, murky water of a fjord in northern Norway.
    orca-mature-male-norway-201601-0276.tif
  • Part of a large group of orcas (Orcinus orca) traveling together while foraging on large schools of herring (Clupea harengus) in the cold waters of northern Norway. With an abundant food supply, most orca social units had at least one calf, as pictured here.
    orcinus-orca-norway-mother-with-calf...tif
  • Swimmer stroking Scar, a male sperm whale who seeks out and enjoys contact with people. Here, he is comfortable enough to have his eyes closed during the encounter. He was around 10 years old at this time.
    scar-sperm-whale-with-snorkeler-domi...tif
  • Natural ice sculpture floating at sea in Svalbard. The bulk of the ice is under the ocean surface.
    ice-sculpture-at-sea-svalbard-201506...tif
  • Bearded seal (Erignathus barbatus) sleeping on ice. The seal is positioned near water so it can slip in quickly if approached, such as by a polar bear.
    bearded-seal-sleeping-erignathus-bar...tif
  • Bearded seal (Erignathus barbatus) resting on ice in Svalbard. Bearded seals are primary prey for polar bears
    bearded-seal-erignathus-barbatus-sva...tif
  • A pair of Arctic terns (Sterna paradisaea) taking a break betweeen periods of fishing. These birds migrate from the Arctic to Antarctica and back each year, spending summer in the northern and southern hemispheres. Round trip distances traveled by these birds involve tens of thousands of kilometers.
    arctic-terns-sterna-paradisaea-svalb...tif
  • A very young polar bear cub (Ursus maritimus), probably only a few months old, standing up to take a look around. This cub was alone with its mother. Given that polar bears generally have two cubs, it is possible that something happened to this cub’s sibling. Photographed in Svalbard.
    polar-bear-cub-ursus-maritumus-svalb...tif
  • A very young polar bear cub (Ursus maritimus), probably only a few months old, trotting away after standing up to take a look around. This cub was alone with its mother. Given that polar bears generally have two cubs, it is possible that something happened to this cub’s sibling. Photographed in Svalbard.
    polar-bear-cub-ursus-maritimus-svalb...tif
  • A very young polar bear cub (Ursus maritimus), probably only a few months old, standing up to take a look around. This cub was alone with its mother. Given that polar bears generally have two cubs, it is possible that something happened to this cub’s sibling. Photographed in Svalbard.
    polar-bear-cub-ursus-maritimus-svalb...tif
  • Polar bear cub (Ursus maritimus) walking on ice in northern Svalbard. This cub was with its mother, playing and groaning loudly to get its mother’s attention.
    polar-bear-cub-ursus-maritimus-svalb...tif
  • Polar bear cub (Ursus maritimus) playing on ice in northern Svalbard.
    polar-bear-cub-on-ice-svalbard-ursus...tif
  • Female polar bear (Ursus maritimus) that had a single young cub, only a few months old. Polar bears usually have two cubs. It is possible that this female lost a cub. The stain on her nose is blood from a seal that she and her cub had been feeding upon. Photographed in northern Svalbard.
    polar-bear-female-ursus-maritimus-sv...tif
  • Four sperm whales (Physeter macrocephalus) rubbing their heads together while socializing, forming an X-pattern. This type of tactile social interaction among sperm whales is common, and is accompanied by a lot of sound made by the whales.
    four-sperm-whales-socializing-x-patt...tif
  • A family unit of sperm whales (Physeter macrocephalus) off the coast of Rouseau, Dominica. The group comprises adult females and juvenile whales. Notice that the individual deepest down is passing gas, which sperm whales seem to do a lot.
    sperm-whale-family-physeter-macrocep...tif
  • Aerial view of a dead humpback whale calf (Megaptera novaeangliae) stranded on a remote beach. This calf was female.
    dead-humpback-whale-calf-south-afric...tif
  • This is a blue ocean slug (Glaucus atlanticus) approaching the venom-filled tentacles of a blue bottle (Physalia utriculus). Both were washed ashore as part of a mass, multi-day stranding of thousands of blue bottles, which are colonial siphonophores that are also called Portuguese man-of-wars. The nudibranchs are part of the rarely seen blue community of the open ocean. They prey on other animals in that community, such as Physalia utriculus, by-the-wind-sailors (Velella velella), blue buttons (Porpita porpita), and violet snails (Janthina janthina). Glaucus nudibranchs are immune to the venom of the blue bottles that they consume. They store their prey’s nematocysts and deploy them in specialized cnidosacs at the tips of their ornate cerata. Because these nudibranchs concentrate the venom of their prey, they can produce a more powerful sting than Physalia.
    glaucus-atlanticus-nudibranch-with-p...tif
  • This is a blue ocean slug (Glaucus atlanticus) that was washed ashore with a mass, multi-day stranding of thousands of blue bottles (Physalia utriculus). These nudibranchs are part of the rarely seen blue community of the open ocean. They prey on other animals in that community, such as Physalia utriculus, by-the-wind-sailors (Velella velella), blue buttons (Porpita porpita), and violet snails (Janthina janthina). Glaucus nudibranchs are immune to the venom of the blue bottles that they consume. They store their prey’s nematocysts and deploy them in specialized cnidosacs at the tips of their ornate cerata. Because these nudibranchs concentrate the venom of their prey, they can produce a more powerful sting than Physalia.
    glaucus-atlanticus-nudibranch-201711...tif
  • Mature bull killer whale (Orcinus orca) in the cold, dark winter waters of northern Norway. Note the rake marks on the orca’s body, perhaps from encounters with other male orcas.
    mature-male-killer-whale-orcinus-orc...tif
  • Mature male killer whale (Orcinus orca) swimming in the low light of winter in northern Norway
    male-killer-whale-orca-low-light-win...tif
  • Three adult southern right whales (Eubalaena australis) engaged in social activity, viewed from the air. Photographed with the permission of the Department of Environmental Affairs, South Africa.
    socializing-southern-right-whales-ae...tif
  • Arctic tern (Sterna paradisaea) in flight in the middle of a beautiful Arctic summer night. This species undertakes an incredible pole to pole migration each year to spend summer in both the northern and the southern hemispheres.
    arctic-tern-flying-overhead-blue-sky...tif
  • Bearded seal (Erignathus barbatus) resting on ice in Svalbard. At full magnification, the seal’s curly whiskers that comprise the beard from which this animal’s common name derives are clearly visible.
    bearded-seal-resting-on-ice-svalbard...tif
  • A very young polar bear cub (Ursus maritimus), probably only a few months old, standing up to take a look around. This cub was alone with its mother. Given that polar bears generally have two cubs, it is possible that something happened to this cub’s sibling. Photographed in Svalbard.
    polar-bear-cub-ursus-maritimus-svalb...tif
  • A very young polar bear cub (Ursus maritimus), probably only a few months old, standing up to take a look around. This cub was alone with its mother. Given that polar bears generally have two cubs, it is possible that something happened to this cub’s sibling. Photographed in Svalbard.
    polar-bear-cub-ursus-maritimus-svalb...tif
  • A very young polar bear cub (Ursus maritimus), probably only a few months old, standing up to take a look around. This cub was alone with its mother. Given that polar bears generally have two cubs, it is possible that something happened to this cub’s sibling. Photographed in Svalbard.
    polar-bear-cub-ursus-maritimus-svalb...tif
  • Social interaction among sperm whales is often a tactile affair. Social gatherings among members of a sperm whale family unit can involve rubbing of bodies against one another and even gentle mouthing, as pictured here. Photo taken under permit in Roseau, Dominica.
    sperm-whales-socializing-dominica-ph...tif
  • A parasitic copepod that was associated with an ocean sunfish (Mola mola). Here, the copepod is backlit and focus-stacked in order to reveal the animal’s inner structure. The long appendage visible at the posterior of the copepod is one of two original appendages. The other appendage fell off during capture. Pictured here, the remaining appendage is split in two, such that the top and bottom halves are visible. The slits in these appendages carried juvenile copepods in various stages of development. This copepod is from the family Caligidae (possibly Caligus sp. or Lepeoptheirus sp.),
    copepod-parasite-mola-sunfish-south-...tif
  • This is an ocean sunfish (Mola mola), around 1.5m in length, basking on the ocean surface near land. One hypothesis concerning why these fish engage in such behavior is that they seek the help of ocean-going birds to remove parasites. Sunfish are host to many species of parasites, including copepods and nematodes.
    mola-mola-ocean-surface-pest-removal...tif
  • Baby sperm whale (Physeter macrocephalus) playing at the ocean surface
    sperm-whale-calf-playing-dominica-20...tif
  • Head-on view of an inquisitive juvenile sperm whale (Physeter macrocephalus) accompanied by large remoras and tiny pilot fish. Note the shedding of the whale's skin, which is common for sperm whales. The position of the whale's single blowhole on the left side of its head is also apparent from this angle of view.
    sperm-whale-frontal-view-physeter-ma...tif
  • This is a small Eutonina indicans hydrozoan, measuring about 4cm in diameter. This species is found throughout the North Pacific and North Atlantic oceans. It inhabits shallow coastal waters.
    eutonina-indicans-hydrozoan-aggregat...tif
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